Spain

Spain
Location
Region	Europe
Subregion	Southern Europe
BioRegion	Palaearctic Region
Diversity
Subfamilies	7
Genera	45
Endemic Genera	0
% Endemic Genera	0.0%
Species/Subspecies	296
Endemic Species	13
% Endemic Species	4.4%
Introduced Species	14

The following valid species and subspecies of ants are known to occur in Spain. Unidentified and undescribed species will be added to this list as identifications are verified and taxonomic revisions are published. This species list is based upon the effort of many ant collectors as well as myrmecologists who have published on the taxonomy of Spanish ants. Those who have specimen records that expand this list are encouraged to submit this information in the discussion section of the Spanish Ants web page. Please advise if any corrections are warranted.

Anochetus

• Anochetus ghilianii (Spinola, 1851)

Aphaenogaster

• Aphaenogaster cardenai Espadaler, 1981
• Aphaenogaster dulcineae Emery, 1924
• Aphaenogaster gemella (Roger, 1862)
• Aphaenogaster gibbosa (Latreille, 1798)
• Aphaenogaster iberica Emery, 1908
• Aphaenogaster ichnusa Santschi, 1925
• Aphaenogaster senilis Mayr, 1853
• Aphaenogaster splendida (Roger, 1859) (introduced)
• Aphaenogaster striativentris Forel, 1895
• Aphaenogaster subterranea (Latreille, 1798)
• Aphaenogaster testaceopilosa (Lucas, 1849)
• Aphaenogaster ulibeli Gomez & Espadaler, 2018

Bothriomyrmex

• Bothriomyrmex corsicus Santschi, 1923
• Bothriomyrmex meridionalis (Roger, 1863)
• Bothriomyrmex saundersi Santschi, 1922

Brachymyrmex

• Brachymyrmex patagonicus Mayr, 1868 (introduced)

Camponotus

• Camponotus aethiops (Latreille, 1798)
• Camponotus amaurus Espadaler, 1997
• Camponotus barbaricus Emery, 1905
• Camponotus cruentatus (Latreille, 1802)
• Camponotus fallax (Nylander, 1856)
• Camponotus foreli Emery, 1881
• Camponotus gestroi Emery, 1878
• Camponotus haroi Espadaler, 1997
• Camponotus herculeanus (Linnaeus, 1758)
• Camponotus honaziensis Karaman & Aktaç, 2013
• Camponotus lateralis (Olivier, 1792)
• Camponotus ligniperda (Latreille, 1802)
• Camponotus micans (Nylander, 1856)
• Camponotus piceus (Leach, 1825)
• Camponotus pilicornis (Roger, 1859)
• Camponotus ruber Emery, 1925
• Camponotus sylvaticus (Olivier, 1792)
• Camponotus universitatis Forel, 1890
• Camponotus vagus (Scopoli, 1763)

Cardiocondyla

• Cardiocondyla batesii Forel, 1894
• Cardiocondyla elegans Emery, 1869
• Cardiocondyla emeryi Forel, 1881
• Cardiocondyla nigra Forel, 1905

Cataglyphis

• Cataglyphis douwesi de Haro & Collingwood, 2000
• Cataglyphis floricola Tinaut, 1993
• Cataglyphis gadeai De Haro & Collingwood, 2003
• Cataglyphis hispanica Emery, 1906
• Cataglyphis humeya Tinaut, 1991
• Cataglyphis iberica Emery, 1906
• Cataglyphis piliscapa (Forel, 1901)
• Cataglyphis rosenhaueri Santschi, 1925
• Cataglyphis tartessica Amor & Ortega, 2014
• Cataglyphis velox Santschi, 1929

Colobopsis

• Colobopsis imitans Schifani, Giannetti, Csősz, Castellucci, Luchetti, Castracani, Spotti, Mori & Grasso, 2021
• Colobopsis truncata (Spinola, 1808)

Crematogaster

• Crematogaster auberti Emery, 1869
• Crematogaster auberti vogti Forel, 1909
• Crematogaster fuentei Menozzi, 1922
• Crematogaster inermis Mayr, 1862
• Crematogaster laestrygon Emery, 1869
• Crematogaster laestrygon submaura Lomnicki, 1925
• Crematogaster scutellaris (Olivier, 1792)
• Crematogaster sordidula (Nylander, 1849)

Cryptopone

• Cryptopone ochracea (Mayr, 1855)

Dolichoderus

• Dolichoderus quadripunctatus (Linnaeus, 1771)

Formica

• Formica bruni Kutter, 1967
• Formica cinerea Mayr, 1853
• Formica clara Forel, 1886
• Formica cunicularia Latreille, 1798
• Formica decipiens Bondroit, 1918
• Formica dusmeti Emery, 1909
• Formica exsecta Nylander, 1846
• Formica foreli Bondroit, 1918
• Formica frontalis Santschi, 1919
• Formica fusca Linnaeus, 1783
• Formica gagates Latreille, 1798
• Formica gerardi Bondroit, 1917
• Formica lemani Bondroit, 1917
• Formica lugubris Zetterstedt, 1838
• Formica picea Nylander, 1846
• Formica polyctena Foerster, 1850
• Formica pratensis Retzius, 1783
• Formica pressilabris Nylander, 1846
• Formica rufa Linnaeus, 1758
• Formica rufibarbis Fabricius, 1793
• Formica sanguinea Latreille, 1798
• Formica selysi Bondroit, 1918

Formicoxenus

• Formicoxenus nitidulus (Nylander, 1846)

Goniomma

• Goniomma baeticum Reyes, Espadaler & Rodriquez, 1987
• Goniomma blanci (André, 1881)
• Goniomma collingwoodi Espadaler, 1997
• Goniomma compressisquama Tinaut, 1995
• Goniomma decipiens Espadaler, 1997
• Goniomma hispanicum André, 1883
• Goniomma kugleri Espadaler, 1986

Harpagoxenus

• Harpagoxenus sublaevis (Nylander, 1849)

Hypoponera

• Hypoponera abeillei (André, 1881)
• Hypoponera eduardi (Forel, 1894)
• Hypoponera nivariana
• Hypoponera punctatissima (Roger, 1859) (introduced)
• Hypoponera ragusai (Emery, 1894)

Iberoformica

• Iberoformica subrufa (Roger, 1859)

Lasius

• Lasius alienus (Foerster, 1850)
• Lasius balearicus Talavera, Espadaler & Vila, 2014
• Lasius bicornis (Foerster, 1850)
• Lasius brunneus (Latreille, 1798)
• Lasius carniolicus Mayr, 1861
• Lasius cinereus Seifert, 1992
• Lasius citrinus Emery, 1922
• Lasius distinguendus (Emery, 1916)
• Lasius emarginatus (Olivier, 1792)
• Lasius flavus (Fabricius, 1782)
• Lasius fuliginosus (Latreille, 1798)
• Lasius grandis Forel, 1909
• Lasius jensi Seifert, 1982
• Lasius lasioides (Emery, 1869)
• Lasius meridionalis (Bondroit, 1920)
• Lasius mixtus (Nylander, 1846)
• Lasius myops Forel, 1894
• Lasius neglectus Van Loon, Boomsma & Andrasfalvy, 1990 (introduced)
• Lasius niger (Linnaeus, 1758)
• Lasius paralienus Seifert, 1992
• Lasius piliferus Seifert, 1992
• Lasius platythorax Seifert, 1991
• Lasius psammophilus Seifert, 1992
• Lasius rabaudi (Bondroit, 1917)
• Lasius sabularum (Bondroit, 1918)
• Lasius turcicus Santschi, 1921 (introduced)
• Lasius umbratus (Nylander, 1846)

Lepisiota

• Lepisiota frauenfeldi (Mayr, 1855)
• Lepisiota nigra (Dalla Torre, 1893)

Leptanilla

• Leptanilla charonea López, Martínez & Barandica, 1994
• Leptanilla plutonia López, Martínez & Barandica, 1994
• Leptanilla revelierii Emery, 1870
• Leptanilla zaballosi López, Martínez & Barandica, 1994

Leptothorax

• Leptothorax acervorum (Fabricius, 1793)
• Leptothorax gredleri Mayr, 1855
• Leptothorax muscorum (Nylander, 1846)

Linepithema

• Linepithema humile (Mayr, 1868) (introduced)

Messor

• Messor barbarus Linnaeus, 1767
• Messor bouvieri Bondroit, 1918
• Messor capitatus (Latreille, 1798)
• Messor celiae Reyes, 1985
• Messor erwini Orou, Csősz, Arnan, Pol, Arthofer, Schlick-Steiner & Steiner, 2023
• Messor hispanicus Santschi, 1919
• Messor ibericus Santschi, 1931
• Messor lobicornis Forel, 1894
• Messor lusitanicus Tinaut, 1985
• Messor marocanus Santschi, 1927
• Messor sordidus Forel, 1892
• Messor structor tyrrhenus Emery, 1898
• Messor timidus Espadaler, 1997

Monomorium

• Monomorium algiricum (Bernard, 1955)
• Monomorium andrei Saunders, 1890 (introduced)
• Monomorium andrei furunculum Emery, 1924
• Monomorium carbonarium (Smith, F., 1858)
• Monomorium monomorium Bolton, 1987
• Monomorium pharaonis (Linnaeus, 1758) (introduced)
• Monomorium salomonis (Linnaeus, 1758) (introduced)
• Monomorium sichelii (Roger, 1862)
• Monomorium subopacum Smith, F., 1858

Myrmecina

• Myrmecina graminicola (Latreille, 1802)

Myrmica

• Myrmica aloba Forel, 1909
• Myrmica babiensis García, Cuesta-Segura & Espadaler, 2024
• Myrmica bibikoffi Kutter, 1963
• Myrmica curvithorax Bondroit, 1920
• Myrmica karavajevi Arnol’di, 1930
• Myrmica lemasnei Bernard, 1967
• Myrmica lobulicornis Nylander, 1857
• Myrmica martini Seifert, Yazdi & Schultz, 2014
• Myrmica rubra (Linnaeus, 17830
• Myrmica ruginodis Nylander, 1846
• Myrmica sabuleti Meinert, 1861
• Myrmica scabrinodis Nylander, 1846
• Myrmica schencki Viereck, 1903
• Myrmica specioides Bondroit, 1918
• Myrmica spinosior Santschi, 1931
• Myrmica sulcinodis Nylander, 1846
• Myrmica vandeli Bondroit, 1920
• Myrmica wesmaeli Bondroit, 1918
• Myrmica xavieri Radchenko, Elmes, & Savolainen, 2008

Nylanderia

• Nylanderia flavipes (Smith, F., 1874) (introduced)
• Nylanderia jaegerskioeldi (Mayr, 1904) (introduced)
• Nylanderia vividula (Nylander, 1846)

Oxyopomyrmex

• Oxyopomyrmex magnus Salata & Borowiec, 2015
• Oxyopomyrmex saulcyi Emery, 1889

Paratrechina

• Paratrechina longicornis (Latreille, 1802) (introduced)

Pheidole

• Pheidole indica Mayr, 1879
• Pheidole pallidula (Nlyander, 1849)

Plagiolepis

• Plagiolepis grassei LeMasne, 1956
• Plagiolepis pallescens Forel, 1889
• Plagiolepis pygmaea (Latreille, 1798)
• Plagiolepis schmitzii Forel, 1895
• Plagiolepis xene Starcke, 1936

Polyergus

• Polyergus rufescens (Latreille, 1798)

Ponera

• Ponera coarctata Latreille, 1802
• Ponera testacea Emery, 1895

Proceratium

• Proceratium melinum (Roger, 1860)

Proformica

• Proformica ferreri Bondroit, 1918
• Proformica longiseta Collingwood, 1978
• Proformica nasuta (Nylander, 1856)

Rossomyrmex

• Rossomyrmex minuchae Tinaut, 1981

Solenopsis

• Solenopsis fairchildi Wheeler, W.M., 1926
• Solenopsis fugax (Latreille, 1798)
• Solenopsis latro Forel, 1894
• Solenopsis lusitanica Emery, 1915
• Solenopsis oraniensis Forel, 1894
• Solenopsis orbula emery, 1875

Stenamma

• Stenamma debile (Foerster, 1850)
• Stenamma striatulum Emery, 1895
• Stenamma westwoodii Westwood, 1839

Stigmatomma

• Stigmatomma emeryi (Saunders, 1890)
• Stigmatomma gaetulicum (Baroni Urbani, 1978)
• Stigmatomma impressifrons Emery, 1869

Strongylognathus

• Strongylognathus caeciliae Forel, 1897
• Strongylognathus huberi Forel, 1874
• Strongylognathus huberi dalmaticus Baroni Urbani, 1969
• Strongylognathus testaceus (Schenck, 1852)

Strumigenys

• Strumigenys argiola (Emery, 1869)
• Strumigenys baudueri Emery, 1875
• Strumigenys membranifera (Emery, 1869)
• Strumigenys tenuipilis (Emery, 1915)

Tapinoma

• Tapinoma darioi Seifert, D'Eustacchio, Kaufmann, Centorame & Modica, 2017
• Tapinoma erraticum (Latreille, 1798)
• Tapinoma ibericum Santschi, 1925
• Tapinoma madeirense Forel, 1895
• Tapinoma magnum Mayr, 1861
• Tapinoma melanocephalum (Fabricius, 1793)
• Tapinoma nigerrimum
• Tapinoma pygmaeum (Dufour, 1857)
• Tapinoma simrothi Krausse-Heldrungen, 1911

Temnothorax

• Temnothorax affinis (Mayr, 1855)
• Temnothorax albipennis (Curtis, 1854)
• Temnothorax alfacarensis Tinaut & Reyes-López, 2020
• Temnothorax algiricus (Forel, 1894)
• Temnothorax angustulus (Nylander, 1856)
• Temnothorax ansei Catarineu, Barberá & Reyes-López, 2017
• Temnothorax atlantis (Santschi, 1911)
• Temnothorax aveli (Bondroit, 1918)
• Temnothorax baeticus (Emery, 1924)
• Temnothorax bejaraniensis Reyes-López & Carpintero-Ortega, 2013
• Temnothorax bernardi (Espadaler, 1982)
• Temnothorax blascoi Espadaler, 1997
• Temnothorax caesari Espadaler, 1997
• Temnothorax cagnianti Tinaut, 1983
• Temnothorax clypeatus (Mayr, 1853)
• Temnothorax continentalis Galkowski & Cagniant, 2017
• Temnothorax corticalis (Schenck, 1852)
• Temnothorax crepuscularis Tinaut, 1995
• Temnothorax cristinae Espadaler, 1997
• Temnothorax curtulus (Santschi, 1929)
• Temnothorax estel González, 2021
• Temnothorax exilis Emery, 1869
• Temnothorax flavispinus (André, 1881)
• Temnothorax formosus (Santschi, 1909)
• Temnothorax gredosi Espadaler & Collingwood, 1982
• Temnothorax grouvellei (Bondroit, 1918)
• Temnothorax ibericus Menozzi, 1922
• Temnothorax interruptus (Schenck, 1852)
• Temnothorax kraussei (Emery, 1915)
• Temnothorax kutteri (Cagniant, 1973)
• Temnothorax leviceps (Emery, 1898)
• Temnothorax lichtensteini (Bondroit, 1918)
• Temnothorax longispinosus (Roger, 1863) (introduced)
• Temnothorax luteus (Forel, 1874)
• Temnothorax mediterraneus Ward et al., 2014
• Temnothorax muellerianus (Finzi, 1922)
• Temnothorax nadigi Kutter, 1925
• Temnothorax naeviventris (Santschi, 1910)
• Temnothorax niger Forel, 1894
• Temnothorax nigriceps (Mayr,, 1855)
• Temnothorax nylanderi (Foerster, 1850)
• Temnothorax pardoi Tinaut, 1987
• Temnothorax parvulus (Schenck, 1852)
• Temnothorax platycephalus (Espadaler, 1997)
• Temnothorax racovitzai (Bondroit, 1918)
• Temnothorax ravouxi (André, 1896)
• Temnothorax recedens (Nylander, 1856)
• Temnothorax schaufussi (Forel, 1879)
• Temnothorax subcingulatus (Emery, 1924)
• Temnothorax tuberum (Fabricius, 175)
• Temnothorax tyndalei (Forel, 1909)
• Temnothorax unifasciatus (Latreille, 1798)
• Temnothorax universitatis (Espadaler, 1997)

Tetramorium

• Tetramorium atratulum (Schenck, 1852)
• Tetramorium bicarinatum (Nylander,, 1846)
• Tetramorium biskrense Forel, 1904
• Tetramorium caespitum (Linnaeus, 1758)
• Tetramorium caldarium (Roger, 1857)
• Tetramorium exasperatum Emery, 1891
• Tetramorium forte Forel, 1904
• Tetramorium impurum (Foerster, 1850)
• Tetramorium indocile Santschi, 1927
• Tetramorium inquilinum Ward et al., 2014
• Tetramorium kutteri (Tinaut, 1990)
• Tetramorium lanuginosum Mayr, 1870
• Tetramorium meridionale Emery, 1870
• Tetramorium pyrenaeicum Roszler, 1937
• Tetramorium semilaeve André, 1883

Wasmannia

• Wasmannia auropunctata (Roger, 1863) (introduced)

References

• Espadaler, X.; García, F.; Roig, X.; Vila, R. 2013. Hormigas (Hymenoptera, Formicidae) del Parc del Castell de Montesquiu (Osona, noreste De La Península Ibérica). Boletín de la Sociedad Entomológica Aragonesa 53:223-227.
• Tinaut, A. 2001. Hypoponera ragusai (Emery, 1895) una hormiga cavernícola nueva para la península Ibérica (Hymenoptera, Formicidae). Graellsia, 57(1), 3-8.
• Introduced species