Taxa Described in 2011

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The following 2 genera were described in 2011 (see all years)

Taxon Name Subfamily Tribe Author Type Species Fossil Fossil Age Species Count Notes
Eldermyrmex Dolichoderinae Shattuck Iridomyrmex oblongiceps, now Eldermyrmex oblongiceps Yes 42–37 Ma Bartonian, Middle to Late Eocene 2
Titanomyrma Formiciinae Archibald, Johnson, Mathewes & Greenwood Titanomyrma gigantea Yes 49.5–47 Ma Ypresian, Early Eocene to Lutetian, Middle Eocene 3

The following 210 species/subspecies were described in 2011 (see all years)

Taxon Name Subfamily Tribe Genus Species Subspecies Author Type Locality Country Fossil Fossil Age Notes
Aenictus bodongjaya Dorylinae Aenictus bodongjaya Jaitrong & Yamane Indonesia
Aenictus brevinodus Dorylinae Aenictus brevinodus Jaitrong & Yamane Indonesia
Aenictus doydeei Dorylinae Aenictus doydeei Jaitrong & Yamane Laos
Aenictus fulvus Dorylinae Aenictus fulvus Jaitrong & Yamane Thailand
Aenictus glabrinotum Dorylinae Aenictus glabrinotum Jaitrong & Yamane Malaysia
Aenictus montivagus Dorylinae Aenictus montivagus Jaitrong & Yamane Malaysia
Aenictus parahuonicus Dorylinae Aenictus parahuonicus Jaitrong & Yamane Thailand
Aenictus rotundicollis Dorylinae Aenictus rotundicollis Jaitrong & Yamane Borneo
Aenictus siamensis Dorylinae Aenictus siamensis Jaitrong & Yamane Thailand
Aenictus sonchaengi Dorylinae Aenictus sonchaengi Jaitrong & Yamane Thailand
Aenictus wayani Dorylinae Aenictus wayani Jaitrong & Yamane Indonesia
Aenictus yamanei Dorylinae Aenictus yamanei Wiwatwitaya & Jaitrong Malaysia
Anochetus maryatiae Ponerinae Ponerini Anochetus maryatiae Nurul Aida & Idris Malaysia
Calyptomyrmex asper Myrmicinae Crematogastrini Calyptomyrmex asper Shattuck Borneo
Calyptomyrmex caledonicus Myrmicinae Crematogastrini Calyptomyrmex caledonicus Shattuck New Caledonia
Calyptomyrmex danum Myrmicinae Crematogastrini Calyptomyrmex danum Shattuck Borneo
Calyptomyrmex fragarus Myrmicinae Crematogastrini Calyptomyrmex fragarus Shattuck Australia
Calyptomyrmex fritillus Myrmicinae Crematogastrini Calyptomyrmex fritillus Shattuck Australia
Calyptomyrmex grammus Myrmicinae Crematogastrini Calyptomyrmex grammus Shattuck Australia
Calyptomyrmex lineolus Myrmicinae Crematogastrini Calyptomyrmex lineolus Shattuck Australia
Calyptomyrmex loweryi Myrmicinae Crematogastrini Calyptomyrmex loweryi Shattuck Philippines
Calyptomyrmex ocullatus Myrmicinae Crematogastrini Calyptomyrmex ocullatus Shattuck Australia
Calyptomyrmex retrostriatus Myrmicinae Crematogastrini Calyptomyrmex retrostriatus Shattuck Borneo
Calyptomyrmex ryderae Myrmicinae Crematogastrini Calyptomyrmex ryderae Shattuck Borneo
Calyptomyrmex sabahensis Myrmicinae Crematogastrini Calyptomyrmex sabahensis Shattuck Borneo
Calyptomyrmex sparsus Myrmicinae Crematogastrini Calyptomyrmex sparsus Shattuck Australia
Calyptomyrmex taylori Myrmicinae Crematogastrini Calyptomyrmex taylori Shattuck Australia
Camponotites steinbachi Formicinae Camponotites steinbachi Dlussky, Karl & Brauckmann Yes Late Pliocene Willershausen, Lower Saxony, Germany
Camponotus iwoensis Formicinae Camponotini Camponotus iwoensis Terayama & Kubota Japan
Carebara abuhurayri Myrmicinae Crematogastrini Carebara abuhurayri Sharaf & Aldawood Synonym, see Carebara arabica
Cataglyphis laylae Formicinae Formicini Cataglyphis laylae Collingwood Synonym, see Cataglyphis saharae
Dolichoderus utriensis Dolichoderinae Dolichoderini Dolichoderus utriensis Ortiz & Fernández Colombia
Dorymyrmex amazonicus Dolichoderinae Leptomyrmecini Dorymyrmex amazonicus Cuezzo & Guerrero Colombia
Dorymyrmex tuberosus Dolichoderinae Leptomyrmecini Dorymyrmex tuberosus Cuezzo & Guerrero Colombia
Dorymyrmex xerophylus Dolichoderinae Leptomyrmecini Dorymyrmex xerophylus Cuezzo & Guerrero Colombia
Heteroponera crozieri Ectatomminae Heteroponerini Heteroponera crozieri Taylor Australia
Heteroponera majeri Ectatomminae Heteroponerini Heteroponera majeri Taylor Australia
Hypoponera aprora Ponerinae Ponerini Hypoponera aprora Bolton & Fisher Tanzania
Hypoponera austra Ponerinae Ponerini Hypoponera austra Bolton & Fisher South Africa
Hypoponera blanda Ponerinae Ponerini Hypoponera blanda Bolton & Fisher Central African Republic
Hypoponera comis Ponerinae Ponerini Hypoponera comis Bolton & Fisher Tanzania
Hypoponera defessa Ponerinae Ponerini Hypoponera defessa Bolton & Fisher Ghana
Hypoponera dema Ponerinae Ponerini Hypoponera dema Bolton & Fisher Rwanda
Hypoponera dis Ponerinae Ponerini Hypoponera dis Bolton & Fisher Tanzania
Hypoponera exigua Ponerinae Ponerini Hypoponera exigua Bolton & Fisher Ethiopia
Hypoponera faex Ponerinae Ponerini Hypoponera faex Bolton & Fisher Ethiopia
Hypoponera fatiga Ponerinae Ponerini Hypoponera fatiga Bolton & Fisher Ghana
Hypoponera hawkesi Ponerinae Ponerini Hypoponera hawkesi Bolton & Fisher Tanzania
Hypoponera hebes Ponerinae Ponerini Hypoponera hebes Bolton & Fisher Rwanda
Hypoponera ignavia Ponerinae Ponerini Hypoponera ignavia Bolton & Fisher Zimbabwe
Hypoponera importuna Ponerinae Ponerini Hypoponera importuna Bolton & Fisher Gabon
Hypoponera jocosa Ponerinae Ponerini Hypoponera jocosa Bolton & Fisher Ethiopia
Hypoponera juxta Ponerinae Ponerini Hypoponera juxta Bolton & Fisher Ethiopia
Hypoponera lassa Ponerinae Ponerini Hypoponera lassa Bolton & Fisher Ghana
Hypoponera lepida Ponerinae Ponerini Hypoponera lepida Bolton & Fisher Ghana
Hypoponera meridia Ponerinae Ponerini Hypoponera meridia Bolton & Fisher South Africa
Hypoponera mixta Ponerinae Ponerini Hypoponera mixta Bolton & Fisher Kenya
Hypoponera molesta Ponerinae Ponerini Hypoponera molesta Bolton & Fisher Gabon
Hypoponera obtunsa Ponerinae Ponerini Hypoponera obtunsa Bolton & Fisher Zimbabwe
Hypoponera odiosa Ponerinae Ponerini Hypoponera odiosa Bolton & Fisher Cameroun
Hypoponera perparva Ponerinae Ponerini Hypoponera perparva Bolton & Fisher South Africa
Hypoponera producta Ponerinae Ponerini Hypoponera producta Bolton & Fisher Gabon
Hypoponera pulchra Ponerinae Ponerini Hypoponera pulchra Bolton & Fisher Equatorial Guinea
Hypoponera quaestio Ponerinae Ponerini Hypoponera quaestio Bolton & Fisher Tanzania
Hypoponera regis Ponerinae Ponerini Hypoponera regis Bolton & Fisher Tanzania
Hypoponera rigida Ponerinae Ponerini Hypoponera rigida Bolton & Fisher Uganda
Hypoponera segnis Ponerinae Ponerini Hypoponera segnis Bolton & Fisher Rwanda
Hypoponera surda Ponerinae Ponerini Hypoponera surda Bolton & Fisher Kenya
Hypoponera tecta Ponerinae Ponerini Hypoponera tecta Bolton & Fisher Kenya
Hypoponera tristis Ponerinae Ponerini Hypoponera tristis Bolton & Fisher Rwanda
Hypoponera venusta Ponerinae Ponerini Hypoponera venusta Bolton & Fisher Rwanda
Iridomyrmex adstringatus Dolichoderinae Leptomyrmecini Iridomyrmex adstringatus Heterick & Shattuck Australia
Iridomyrmex alpinus Dolichoderinae Leptomyrmecini Iridomyrmex alpinus Heterick & Shattuck Australia
Iridomyrmex atypicus Dolichoderinae Leptomyrmecini Iridomyrmex atypicus Heterick & Shattuck Australia
Iridomyrmex brennani Dolichoderinae Leptomyrmecini Iridomyrmex brennani Heterick & Shattuck Australia
Iridomyrmex coeruleus Dolichoderinae Leptomyrmecini Iridomyrmex coeruleus Heterick & Shattuck Australia
Iridomyrmex cuneiceps Dolichoderinae Leptomyrmecini Iridomyrmex cuneiceps Heterick & Shattuck Australia
Iridomyrmex cupreus Dolichoderinae Leptomyrmecini Iridomyrmex cupreus Heterick & Shattuck Australia
Iridomyrmex curvifrons Dolichoderinae Leptomyrmecini Iridomyrmex curvifrons Heterick & Shattuck Australia
Iridomyrmex difficilis Dolichoderinae Leptomyrmecini Iridomyrmex difficilis Heterick & Shattuck Australia
Iridomyrmex elongatus Dolichoderinae Leptomyrmecini Iridomyrmex elongatus Heterick & Shattuck Australia
Iridomyrmex fulgens Dolichoderinae Leptomyrmecini Iridomyrmex fulgens Heterick & Shattuck Australia
Iridomyrmex gibbus Dolichoderinae Leptomyrmecini Iridomyrmex gibbus Heterick & Shattuck Australia
Iridomyrmex gumnos Dolichoderinae Leptomyrmecini Iridomyrmex gumnos Heterick & Shattuck Australia
Iridomyrmex hertogi Dolichoderinae Leptomyrmecini Iridomyrmex hertogi Heterick & Shattuck Australia
Iridomyrmex infuscus Dolichoderinae Leptomyrmecini Iridomyrmex infuscus Heterick & Shattuck Australia
Iridomyrmex longisoma Dolichoderinae Leptomyrmecini Iridomyrmex longisoma Heterick & Shattuck Australia
Iridomyrmex luteoclypeatus Dolichoderinae Leptomyrmecini Iridomyrmex luteoclypeatus Heterick & Shattuck Australia
Iridomyrmex macrops Dolichoderinae Leptomyrmecini Iridomyrmex macrops Heterick & Shattuck Australia
Iridomyrmex meridianus Dolichoderinae Leptomyrmecini Iridomyrmex meridianus Heterick & Shattuck Australia
Iridomyrmex mirabilis Dolichoderinae Leptomyrmecini Iridomyrmex mirabilis Heterick & Shattuck Australia
Iridomyrmex neocaledonica Dolichoderinae Leptomyrmecini Iridomyrmex neocaledonica Heterick & Shattuck New Caledonia
Iridomyrmex niger Dolichoderinae Leptomyrmecini Iridomyrmex niger Heterick & Shattuck Australia
Iridomyrmex nudipes Dolichoderinae Leptomyrmecini Iridomyrmex nudipes Heterick & Shattuck Australia
Iridomyrmex omalonotus Dolichoderinae Leptomyrmecini Iridomyrmex omalonotus Heterick & Shattuck Australia
Iridomyrmex phillipensis Dolichoderinae Leptomyrmecini Iridomyrmex phillipensis Heterick & Shattuck Australia
Iridomyrmex roseatus Dolichoderinae Leptomyrmecini Iridomyrmex roseatus Heterick & Shattuck Australia
Iridomyrmex suchieroides Dolichoderinae Leptomyrmecini Iridomyrmex suchieroides Heterick & Shattuck Australia
Iridomyrmex tenebrans Dolichoderinae Leptomyrmecini Iridomyrmex tenebrans Heterick & Shattuck Australia
Iridomyrmex tenuiceps Dolichoderinae Leptomyrmecini Iridomyrmex tenuiceps Heterick & Shattuck Australia
Iridomyrmex trigonoceps Dolichoderinae Leptomyrmecini Iridomyrmex trigonoceps Heterick & Shattuck Australia
Iridomyrmex xanthocoxa Dolichoderinae Leptomyrmecini Iridomyrmex xanthocoxa Heterick & Shattuck Australia
Lepisiota elegantissima Formicinae Plagiolepidini Lepisiota elegantissima Collingwood & van Harten United Arab Emirates
Leptanilloides erinys Dorylinae Leptanilloides erinys Borowiec & Longino Ecuador
Leptanilloides femoralis Dorylinae Leptanilloides femoralis Borowiec & Longino Venezuela
Leptanilloides gracilis Dorylinae Leptanilloides gracilis Borowiec & Longino Mexico
Leptogenys amu Ponerinae Ponerini Leptogenys amu Lattke Colombia
Leptogenys bifida Ponerinae Ponerini Leptogenys bifida Lattke Honduras
Leptogenys carbonaria Ponerinae Ponerini Leptogenys carbonaria Lattke Colombia
Leptogenys chamela Ponerinae Ponerini Leptogenys chamela Lattke Mexico
Leptogenys ciliata Ponerinae Ponerini Leptogenys ciliata Lattke Ecuador
Leptogenys cordoba Ponerinae Ponerini Leptogenys cordoba Lattke Mexico
Leptogenys corniculans Ponerinae Ponerini Leptogenys corniculans Lattke Brazil
Leptogenys cracens Ponerinae Ponerini Leptogenys cracens Lattke Mexico
Leptogenys cuneata Ponerinae Ponerini Leptogenys cuneata Lattke Ecuador
Leptogenys deborae Ponerinae Ponerini Leptogenys deborae Lattke Venezuela
Leptogenys erugata Ponerinae Ponerini Leptogenys erugata Lattke Venezuela
Leptogenys foraminosa Ponerinae Ponerini Leptogenys foraminosa Lattke Costa Rica
Leptogenys foveonates Ponerinae Ponerini Leptogenys foveonates Lattke Venezuela
Leptogenys gatu Ponerinae Ponerini Leptogenys gatu Lattke Panama
Leptogenys glabra Ponerinae Ponerini Leptogenys glabra Lattke Colombia
Leptogenys gorgona Ponerinae Ponerini Leptogenys gorgona Lattke Colombia
Leptogenys ixta Ponerinae Ponerini Leptogenys ixta Lattke Mexico
Leptogenys kiche Ponerinae Ponerini Leptogenys kiche Lattke Guatemala
Leptogenys linda Ponerinae Ponerini Leptogenys linda Lattke Colomblia
Leptogenys mavaca Ponerinae Ponerini Leptogenys mavaca Lattke Venezuela
Leptogenys maya Ponerinae Ponerini Leptogenys maya Lattke Mexico
Leptogenys melena Ponerinae Ponerini Leptogenys melena Lattke Venezuela
Leptogenys minima Ponerinae Ponerini Leptogenys minima Lattke Brazil
Leptogenys montuosa Ponerinae Ponerini Leptogenys montuosa Lattke Panama
Leptogenys nigricans Ponerinae Ponerini Leptogenys nigricans Lattke Brazil
Leptogenys oaxaca Ponerinae Ponerini Leptogenys oaxaca Lattke Mexico
Leptogenys orchidioides Ponerinae Ponerini Leptogenys orchidioides Lattke Guatemala
Leptogenys panops Ponerinae Ponerini Leptogenys panops Lattke French Guiana
Leptogenys paraensis Ponerinae Ponerini Leptogenys paraensis Lattke Brazil
Leptogenys peruana Ponerinae Ponerini Leptogenys peruana Lattke Peru
Leptogenys phylloba Ponerinae Ponerini Leptogenys phylloba Lattke Colombia
Leptogenys pinna Ponerinae Ponerini Leptogenys pinna Lattke Costa Rica
Leptogenys pittieri Ponerinae Ponerini Leptogenys pittieri Lattke Venezuela
Leptogenys pucuna Ponerinae Ponerini Leptogenys pucuna Lattke Ecuador
Leptogenys quadrata Ponerinae Ponerini Leptogenys quadrata Lattke Colombia
Leptogenys quirozi Ponerinae Ponerini Leptogenys quirozi Lattke Mexico
Leptogenys rasila Ponerinae Ponerini Leptogenys rasila Lattke Ecuador
Leptogenys reggae Ponerinae Ponerini Leptogenys reggae Lattke Jamaica
Leptogenys santacruzi Ponerinae Ponerini Leptogenys santacruzi Lattke Ecuador
Leptogenys serrata Ponerinae Ponerini Leptogenys serrata Lattke Panama
Leptogenys sianka Ponerinae Ponerini Leptogenys sianka Lattke Mexico
Leptogenys socorda Ponerinae Ponerini Leptogenys socorda Lattke Colombia
Leptogenys sonora Ponerinae Ponerini Leptogenys sonora Lattke Mexico
Leptogenys tama Ponerinae Ponerini Leptogenys tama Lattke Colombia
Leptogenys tiobil Ponerinae Ponerini Leptogenys tiobil Lattke Venezuela
Leptogenys toxeres Ponerinae Ponerini Leptogenys toxeres Lattke Costa Rica
Leptogenys volcanica Ponerinae Ponerini Leptogenys volcanica Lattke Costa Rica
Leptogenys yocota Ponerinae Ponerini Leptogenys yocota Lattke Honduras
Leptothorax zhengi Myrmicinae Crematogastrini Leptothorax zhengi Zhou & Chen Synonym, see Formicoxenus sibiricus
Monomorium dryhimi Myrmicinae Solenopsidini Monomorium dryhimi Aldawood & Sharaf Synonym, see Monomorium exiguum
Myrmica elmesi Myrmicinae Myrmicini Myrmica elmesi Bharti & Sharma India
Myrmica longisculpta Myrmicinae Myrmicini Myrmica longisculpta Bharti & Sharma India
Myrmica radchenkoi Myrmicinae Myrmicini Myrmica radchenkoi Bharti & Sharma India
Myrmica zhengi Myrmicinae Myrmicini Myrmica zhengi Ma & Xu, S. Synonym, see Myrmica luteola
Myrmoteras glabrum Formicinae Myrmoteratini Myrmoteras glabrum Zettel & Sorger Philippines
Myrmoteras mcarthuri Formicinae Myrmoteratini Myrmoteras mcarthuri Zettel & Sorger Philippines
Neivamyrmex albacorpus Dorylinae Neivamyrmex albacorpus Varela-Hernández & Castaño-Meneses Mexico
Nylanderia boltoni Formicinae Lasiini Nylanderia boltoni LaPolla & Fisher Gabon
Nylanderia brevisetula Formicinae Lasiini Nylanderia brevisetula LaPolla & Fisher Gabon
Nylanderia impolita Formicinae Lasiini Nylanderia impolita LaPolla & Fisher Gabon
Nylanderia luteafra Formicinae Lasiini Nylanderia luteafra LaPolla & Fisher Gabon
Nylanderia scintilla Formicinae Lasiini Nylanderia scintilla LaPolla & Fisher Ivory Coast
Nylanderia silvula Formicinae Lasiini Nylanderia silvula LaPolla & Fisher Kenya
Nylanderia umbella Formicinae Lasiini Nylanderia umbella LaPolla & Fisher Cameroun
Nylanderia usambarica Formicinae Lasiini Nylanderia usambarica LaPolla, Hawkes & Fisher Tanzania
Odontomachus alius Ponerinae Ponerini Odontomachus alius Sorger & Zettel Philippines
Odontomachus schoedli Ponerinae Ponerini Odontomachus schoedli Sorger & Zettel Philippines
Odontomachus scifictus Ponerinae Ponerini Odontomachus scifictus Sorger & Zettel Philippines
Oxyepoecus bidentatus Myrmicinae Solenopsidini Oxyepoecus bidentatus Delsinne & Mackay Paraguay
Oxyepoecus striatus Myrmicinae Solenopsidini Oxyepoecus striatus Mackay, W.P. & Delsinne Paraguay
Paratopula zhengi Myrmicinae Crematogastrini Paratopula zhengi Xu, Z. & Xu, G. China
Pheidole acutilobata Myrmicinae Attini Pheidole acutilobata Mackay, Sola & Josens Argentina
Pheidole hainanensis Myrmicinae Attini Pheidole hainanensis Chen, Ye, Lu & Zhou China
Pheidole iceni Myrmicinae Attini Pheidole iceni Fernández Colombia
Pheidole rasnitsyni Myrmicinae Attini Pheidole rasnitsyni Dubovikoff Baltic Sea area
Plagiolepis boltoni Formicinae Plagiolepidini Plagiolepis boltoni Sharaf & Aldawood Saudi Arabia
Plagiolepis juddi Formicinae Plagiolepidini Plagiolepis juddi Sharaf & Aldawood Yemen
Platythyrea dlusskyi Ponerinae Platythyreini Platythyrea dlusskyi Aria, Perrichot & Nel Yes Ypresian, Early Eocene Oise amber, France
Pristomyrmex rasnitsyni Myrmicinae Crematogastrini Pristomyrmex rasnitsyni Dlussky & Radchenko Yes Bartonian, Middle to Late Eocene Danish-Scandinavian amber, Denmark
Protanilla boltoni Leptanillinae Leptanillini Protanilla boltoni Borowiec, Schulz, Alpert & Banas Malaysia
Protanilla helenae Leptanillinae Leptanillini Protanilla helenae Borowiec, Schulz, Alpert & Banas Philippines
Solenopsis saudiensis Myrmicinae Solenopsidini Solenopsis saudiensis Sharaf & Aldawood Synonym, see Solenopsis abdita
Sphinctomyrmex marcoyi Dorylinae Sphinctomyrmex marcoyi Feitosa, Brandão, Fernández & Delabie Brazil
Sphinctomyrmex schoerederi Dorylinae Sphinctomyrmex schoerederi Feitosa, Brandão, Fernández & Delabie Brazil
Stenamma ailaoense Myrmicinae Stenammini Stenamma ailaoense Liu & Xu China
Stenamma siculum Myrmicinae Stenammini Stenamma siculum Rigato Italy
Stenamma wumengense Myrmicinae Stenammini Stenamma wumengense Liu & Xu China
Stenamma yaluzangbum Myrmicinae Stenammini Stenamma yaluzangbum Liu & Xu China
Stenamma zanoni Myrmicinae Stenammini Stenamma zanoni Rigato Italy
Stigmatomma boltoni Amblyoponinae Amblyoponini Stigmatomma boltoni Bharti & Wachkoo India
Strongylognathus kabakovi Myrmicinae Crematogastrini Strongylognathus kabakovi Radchenko & Dubovikoff Afghanistan
Strumigenys nankunshana Myrmicinae Attini Strumigenys nankunshana Zhou China
Technomyrmex montaseri Dolichoderinae Technomyrmex montaseri Sharaf, Collingwood & Aldawood Oman
Temnothorax liviae Myrmicinae Crematogastrini Temnothorax liviae Agosti & Collingwood United Arab Emirates
Temnothorax mekira Myrmicinae Crematogastrini Temnothorax mekira Terayama & Kubota Japan
Tetramorium azcatltontlium Myrmicinae Crematogastrini Tetramorium azcatltontlium Marques, Vásquez-Bolaños & Quesada Mexico
Tetramorium hasinae Myrmicinae Crematogastrini Tetramorium hasinae Yamane & Jaitrong Thailand
Tetramorium lucyae Myrmicinae Crematogastrini Tetramorium lucyae Sorger Malaysia
Tetramorium mahafaly Myrmicinae Crematogastrini Tetramorium mahafaly Hita Garcia & Fisher Madagascar
Tetramorium meghalayense Myrmicinae Crematogastrini Tetramorium meghalayense Bharti India
Tetramorium notomelanum Myrmicinae Crematogastrini Tetramorium notomelanum Vásquez-Bolaños, Castaño-Meneses & Guzmán-Mendoza Mexico
Tetramorium polymorphum Myrmicinae Crematogastrini Tetramorium polymorphum Yamane & Jaitrong Laos
Titanomyrma lubei Formiciinae Titanomyrma lubei Archibald, Johnson, Mathewes & Greenwood Yes Ypresian, Early Eocene
Green River Formation, Sweetwater County, Wyoming, United States
Turneria rosschinga Dolichoderinae Turneria rosschinga Shattuck Australia