Checklist of Dorylus species

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Dorylus
Dorylinae
Dorylus helvolus casent0106079 head 1.jpg
Dorylus helvolus
Dorylus Distribution.png
Distribution
Dorylus Species Richness.png
Species Richness
Statistics
Valid Species 127
Invalid Species 40
Fossils 0
Countries Occupied 60
Year Described 1793
First Species Described 1764
Last Species Described 1975

The following species and subspecies belong to the genus Dorylus. Synonyms are listed under their senior names. For valid names only see Dorylus species and for distribution information see Dorylus species by Country.

A

acutus Santschi, 1937 (Democratic Republic of Congo)
aethiopicus Emery, 1895 (Eritrea, Guinea, Ivory Coast, Mali, Sudan)
affinis Shuckard, 1840 (Chad, Gambia, Ghana, Guinea, Guinea-Bissau, Ivory Coast, Kenya, Liberia, Malawi, Mali, Senegal, Turkey, Zimbabwe)
   • Dorylus planiceps Haldeman, 1849
   • Typhlopone oraniensis brevinodosus Mayr, 1862

affinis aegyptiacus Mayr, 1865 (Egypt, Eritrea, Ethiopia)
   • Dorylus brevinodosus abyssinicus Emery, 1895

affinis denudatus Santschi, 1910 (Niger)
affinis exilis Santschi, 1914 (Kenya, United Republic of Tanzania)
affinis hirsutus Wheeler, W.M., 1922 (Egypt, Ethiopia)
affinis loewyi Forel, 1907 (Eritrea, United Republic of Tanzania)
affinis parapsidalis Santschi, 1917 (Malawi)
affinis pulliceps Santschi, 1917 (Ivory Coast)
aggressor Santschi, 1923 (Democratic Republic of Congo)
alluaudi Santschi, 1914 (Uganda)
alluaudi lobatus Santschi, 1919 (Democratic Republic of Congo)
atratus Smith, F., 1859 (Nigeria)
atriceps Shuckard, 1840 (Gambia, Guinea, Ivory Coast, Kenya)
   • Dorylus ritsemae Dalla Torre, 1892
   • Dorylus shuckardi Ritsema, 1874

attenuatus Shuckard, 1840 (Botswana, Gambia, Namibia)
   • Dorylus attenuatus umbratipennis Forel, 1909

attenuatus acuminatus Emery, 1899 (South Africa)
attenuatus australis Santschi, 1919 (South Africa)
attenuatus bondroiti Santschi, 1912 (South Africa)
attenuatus latinodis Forel, 1920 (Democratic Republic of Congo)

B

bequaerti Forel, 1913 (Democratic Republic of Congo, Mozambique, South Africa)
bishyiganus (Boven, 1972) (Rwanda)
braunsi Emery, 1895 (Equatorial Guinea, Kenya, Liberia, Zimbabwe)
   • Dorylus helvolus impressus Stitz, 1910

braunsi anceps Forel, 1914 (Zimbabwe)
brevipennis Emery, 1895 (Guinea, Ivory Coast, Kenya, United Republic of Tanzania, Zimbabwe)
brevipennis marshalli Emery, 1901 (Zimbabwe)
brevipennis zimmermanni Santschi, 1910 (Congo)
brevis Santschi, 1919 (Democratic Republic of Congo)
buyssoni Santschi, 1910 (Kenya)
buyssoni conjugens Santschi, 1910 (Kenya)

C

congolensis Santschi, 1910 (Congo)
conradti Emery, 1895 (Ivory Coast, Kenya, Togo, Uganda)
conradti berlandi Santschi, 1926 (Ivory Coast)

D

depilis Emery, 1895 (Cameroun, Kenya, Zimbabwe)
depilis clarior Santschi, 1917 (Democratic Republic of Congo)
depilis ugandensis Santschi, 1914 (Uganda)
diadema Gerstäcker, 1859 (Malawi, Mozambique, Zimbabwe)
diadema arnoldi Forel, 1914 (Zimbabwe)
diadema fusciceps Emery, 1899 (Malawi)
distinctus Santschi, 1910 (Guinea)
ductor Santschi, 1939 (Congo)

E

emeryi Mayr, 1896 (Cameroun, Central African Republic)
emeryi opacus Forel, 1909 (Cameroun, Central African Republic, Democratic Republic of Congo, Guinea-Bissau)
emeryi pulsi (Forel, 1904)
erraticus (Smith, F., 1865) (Guinea-Bissau)

F

faurei Arnold, 1946 (South Africa)
fimbriatus (Shuckard, 1840) (Gambia, Guinea, Kenya, Mali, Sierra Leone, Zimbabwe)
fimbriatus crampeli Santschi, 1919 (Congo)
fimbriatus laevipodex Santschi, 1919 (Kenya)
fimbriatus poweri Forel, 1914 (South Africa)
fulvus (Westwood, 1839) (Algeria, Balearic Islands, Central African Republic, Gambia, Guinea, India, Ivory Coast, Kenya, Mozambique, Namibia, Oman, Pakistan, Saudi Arabia, Senegal, Sierra Leone, Turkey, United Republic of Tanzania, Zimbabwe)
   • Cosmaecetes homalinus Spinola, 1851
   • Labidus kirbii Shuckard, 1840
   • Labidus spinolae Shuckard Shuckard, 1840
   • Labidus thwaitsii Shuckard, 1840
   • Typhlopone clausii Joseph, 1882
   • Typhlopone dahlbomii Westwood, 1840
   • Typhlopone shuckardi Westwood, 1840

fulvus badius Gerstäcker, 1859 (Kenya, Mozambique, Zimbabwe)
   • Dorylus fulvus rhodesiae Forel, 1913

fulvus crosi Santschi, 1926 (Algeria)
fulvus dentifrons Wasmann, 1904 (Cameroun, Democratic Republic of Congo)
   • Dorylus fulvus stramineus Stitz, 1910

fulvus eurous Emery, 1915 (Eritrea, Ethiopia)
fulvus glabratus Shuckard, 1840 (Gambia)
fulvus juvenculus Shuckard, 1840 (Algeria, India, Morocco, Turkey)
   • Typhlopone oraniensis Lucas, 1849

fulvus mordax Santschi, 1931 (Ivory Coast)
   • Dorylus fulvus impressus Santschi, 1928

fulvus obscurior Wheeler, W.M., 1925 (Guinea)
fulvus punicus Santschi, 1926 (Israel, Tunisia)
fulvus ruficeps Santschi, 1926 (Israel, Lebanon)
fulvus saharensis Santschi, 1926
funereus Emery, 1895 (Ghana, Guinea-Bissau)
funereus acherontus Santschi, 1937 (Congo)
funereus pardus Santschi, 1937 (Democratic Republic of Congo)
funereus stygis Santschi, 1937 (Democratic Republic of Congo)
funereus zumpti Santschi, 1937 (Cameroun)
furcatus (Gerstäcker, 1872) (South Africa)
fuscipennis (Emery, 1892) (Ghana, Ivory Coast)
fuscipennis lugubris Santschi, 1919 (Ivory Coast)
fuscipennis marginiventris Santschi, 1919 (Ivory Coast)

G

gaudens Santschi, 1919 (Democratic Republic of Congo)
ghanensis Boven, 1975 (Ghana)
gribodoi Emery, 1892 (Ghana, Guinea, Guinea-Bissau, Ivory Coast, Sierra Leone, Togo)
   • Dorylus gerstaeckeri Emery, 1895
   • Dorylus gribodoi confusa Santschi, 1915
   • Dorylus gribodoi insularis Santschi, 1937
   • Dorylus lamottei Bernard, 1953

H

helvolus (Linnaeus, 1764) (Italy, Kenya, Malawi, Mozambique, South Africa, United Republic of Tanzania)
   • Dorylus planifrons Mayr, 1865
   • Mutilla dorylus Lamarck, 1817
   • Typhlopone europaea Roger, 1859
   • Typhlopone punctata Smith, F., 1858

helvolus pretoriae Arnold, 1946 (South Africa)

K

katanensis Stitz, 1911 (Democratic Republic of Congo)
kohli Wasmann, 1904 (Cameroun, Democratic Republic of Congo, Guinea-Bissau, Kenya, Uganda, United Republic of Tanzania)
   • Dorylus gerstaeckeri quadratus Santschi, 1914

kohli chapini Wheeler, W.M., 1922 (Democratic Republic of Congo)
kohli frenisyi Forel, 1916 (Democratic Republic of Congo)
kohli indocilis Santschi, 1933 (Democratic Republic of Congo)
kohli langi Wheeler, W.M., 1922 (Democratic Republic of Congo)
kohli militaris Santschi, 1923 (Democratic Republic of Congo, Guinea-Bissau)
kohli minor Santschi, 1911 (Angola)
kohli victoriae Santschi, 1921 (Guinea, Uganda)

L

labiatus Shuckard, 1840 (Bangladesh, China, India, Nepal, Pakistan, Sri Lanka)
   • Dorylus hindostanus Smith, F., 1859
   • Dorylus laeviceps Smith, F., 1878

laevigatus (Smith, F., 1857) (Borneo, China, India, Indonesia, Laos, Malaysia, Myanmar, Thailand, Vietnam)
   • Dichthadia glaberrima Gerstäcker, 1863
   • Dorylus breviceps Emery, 1889
   • Dorylus klugi Emery, 1887

leo Santschi, 1919 (Ivory Coast, Mali)

M

mandibularis Mayr, 1896 (Cameroun, Ivory Coast)
mandibularis pulchellus Santschi, 1920 (Ivory Coast)
mayri Santschi, 1912 (Cameroun)
moestus Emery, 1895 (Democratic Republic of Congo, Guinea, Ivory Coast, Kenya, Liberia)
moestus claripennis Santschi, 1919 (Democratic Republic of Congo)
moestus morio Santschi, 1919 (Congo)
moestus schereri Forel, 1911 (Liberia)
montanus Santschi, 1910 (United Republic of Tanzania)

N

niarembensis (Boven, 1972) (Democratic Republic of Congo)
nigricans Illiger, 1802 (Guinea-Bissau, Ivory Coast, Liberia, Sierra Leone, United Republic of Tanzania)
nigricans arcens (Westwood, 1847) (Guinea, Ivory Coast, Liberia, Sierra Leone)
   • Anomma pubescens Roger, 1861

nigricans burmeisteri (Shuckard, 1840) (Benin, Guinea, Ivory Coast, Kenya, Liberia, Mali, Mozambique, Senegal, Sierra Leone, Zimbabwe)
nigricans molestus (Gerstäcker, 1859) (Ethiopia, Guinea-Bissau, Kenya, Mozambique, United Republic of Tanzania, Zimbabwe)
   • Alaopone antinorii Emery, 1881

nigricans pallidus Santschi, 1921 (Cameroun)
nigricans rubellus (Savage, 1849)
nigricans sjostedti Emery, 1899 (Cameroun, Central African Republic, Chad)
nigricans sjostedtiwilverthi (Wasmann, 1917) (Cameroun)
nigricans terrificus Santschi, 1923 (Democratic Republic of Congo, Guinea, Ivory Coast, Uganda)

O

ocellatus (Stitz, 1910) (Cameroun)
orientalis Westwood, 1835 (Bangladesh, Borneo, China, India, Indonesia, Laos, Malaysia, Myanmar, Nepal, Pakistan, Sri Lanka, Thailand, Vietnam)
   • Alaopone oberthueri Emery, 1881
   • Dorylus fuscus Emery, 1889
   • Dorylus longicornis Shuckard, 1840
   • Labidus curtisii Shuckard, 1840

orientalis obscuriceps Santschi, 1920 (India)

P

politus Emery, 1901 (Cameroun)

R

rufescens Santschi, 1915 (Cameroun, Gambia, Senegal)

S

savagei Emery, 1895 (Gabon, Guinea)
savagei mucronatus Emery, 1899 (Nigeria)
schoutedeni Santschi, 1923 (Democratic Republic of Congo)
spininodis Emery, 1901 (Cameroun, Guinea)
spininodis longiceps Viehmeyer, 1914 (United Republic of Tanzania)
stadelmanni Emery, 1895 (Democratic Republic of Congo)
stanleyi Forel, 1909 (Democratic Republic of Congo, Kenya)
staudingeri Emery, 1895 (Democratic Republic of Congo)
striatidens Santschi, 1910 (Ethiopia, Senegal, Zambia)
sudanicus Santschi, 1917 (Chad, Congo, Gambia, Mali)

T

termitarius Wasmann, 1911 (Congo, Democratic Republic of Congo)
titan Santschi, 1923 (Democratic Republic of Congo, Guinea)
titan vinalli Santschi, 1933 (Democratic Republic of Congo)

V

vishnui Wheeler, W.M., 1913 (China, Myanmar, Thailand)

W

westwoodii (Shuckard, 1840)
wilverthi Emery, 1899 (Cameroun, Central African Republic, Democratic Republic of Congo, Ivory Coast, Kenya, Uganda)
   • Dorylus nigritarsis Strand, 1911
   • Dorylus nomadas Santschi, 1935