Checklist of Formica species

AntWiki: The Ants --- Online
Jump to navigation Jump to search
  This is an AntWiki Report  
Formica
Formicinae
Formica rufa casent0173862 head 1.jpg
Formica rufa
Formica Distribution.png
Distribution
Formica Species Richness.png
Species Richness
Statistics
Valid Species 280
Invalid Species 163
Fossils 68
Countries Occupied 76
Year Described 1758
First Species Described 1758
Last Species Described 2009

The following species and subspecies belong to the genus Formica. Synonyms are listed under their senior names. For valid names only see Formica species and for distribution information see Formica species by Country.

A

abdominalis Latreille, 1802
accreta Francoeur, 1973 (Canada, United States)
adamsi Wheeler, W.M., 1909 (Canada, United States)
adamsi alpina Wheeler, W.M., 1909 (United States)
adamsi whymperi Wheeler, W.M., 1917 (Canada, United States)
   • Formica microgyna californica Wheeler, W.M., 1917

adelungi Forel, 1904 (Mongolia)
aegyptiaca Fabricius, 1775
aequalis Walker, 1871
aerata Francoeur, 1973 (United States)
affinis Leach, 1825
albipennis Fabricius, 1793
altayensis Xia & Zheng, 1997 (China)
altipetens Wheeler, W.M., 1913 (Canada, Mexico, United States)
amyoti Le Guillou, 1842
anatolica Seifert & Schultz, 2009 (Turkey)
animosa Forskål, 1775
approximans Wheeler, W.M., 1933 (China)
aquilonia Yarrow, 1955 (Austria, Belarus, Bulgaria, China, Czech Republic, Democratic Peoples Republic of Korea, Denmark, Estonia, Finland, France, Georgia, Germany, Italy, Latvia, Lithuania, Mongolia, Montenegro, Norway, Poland, Romania, Russian Federation, Serbia, Slovenia, Sweden, Switzerland, Ukraine, United Kingdom of Great Britain and Northern Ireland)
   • Formica rufa schmidti Ruzsky, 1926

archboldi Smith, M.R., 1944 (United States)
arenicola Buckley, 1866
argentea Wheeler, W.M., 1912 (Canada, Mexico, United States)
   • Formica fusca argentata Wheeler, W.M., 1902
   • Formica fusca blanda Wheeler, W.M., 1913

aserva Forel, 1901 (Canada, United States)
   • Formica sanguinea subnuda Wheeler, W.M., 1910

aseta Chang & He, 2002 (China)
aterrima Cresson, 1865 (United States)
atra Schilling Schilling, 1839

B

beijingensis Wu, J., 1990 (China)
bicolor Leach, 1825
biophilica Trager, 2007 (United States)
bradleyi Wheeler, W.M., 1913 (Canada, United States)
   • Formica neogagates morbida Wheeler, W.M., 1913

breviscapa Chang & He, 2002 (China)
browni Francoeur, 1973 (Mexico)
bruni Kutter, 1967 (Austria, Germany, Greece, Iberian Peninsula, Spain, Sweden, Switzerland)

C

californica (Creighton, 1950) (United States)
calviceps Cole, 1954 (Mexico, United States)
canadensis Santschi, 1914 (Canada, United States)
candida Smith, F., 1878 (Albania, Armenia, Austria, Belarus, Belgium, China, Democratic Peoples Republic of Korea, Denmark, Estonia, Finland, Georgia, Germany, India, Japan, Kyrgyzstan, Latvia, Lithuania, Luxembourg, Mongolia, Netherlands, Norway, Poland, Republic of Korea, Romania, Russian Federation, Slovakia, Sweden, Switzerland, Ukraine, United Kingdom of Great Britain and Northern Ireland)
candida formosae Emery, 1925 (Taiwan)
chufejif Forskål, 1775
ciliata Mayr, 1886 (United States)
cinerea Mayr, 1853 (Albania, Armenia, Austria, Belarus, Bosnia and Herzegovina, Bulgaria, China, Croatia, Czech Republic, Denmark, Estonia, Finland, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Iran, Italy, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine)
   • Formica balcanina Petrov & Collingwood, 1993
   • Formica cinerea armenica Ruzsky, 1905
   • Formica cinerea brevisetosa Karavaiev, 1927
   • Formica cinerea iberica Finzi, 1928
   • Formica cinerea imitans Ruzsky, 1902
   • Formica cinerea italica Finzi, 1928
   • Formica cinerea novaki Kratochvíl, 1941
   • Formica cinerea ochracea Karavaiev, 1937
   • Formica cinerea sabulosa Karavaiev, 1931
   • Formica cinerea subrufoides Forel, 1913

cinerea cinereoimitans Ruzsky, 1905 (Russian Federation)
cinereofusca Karavaiev, 1929 (Armenia, China, Georgia, Russian Federation, Ukraine)
clara Forel, 1886 (Afghanistan, Austria, Azerbaijan, Belgium, Bulgaria, China, Croatia, Czech Republic, Georgia, Germany, Greece, Hungary, India, Iran, Israel, Italy, Kyrgyzstan, Mongolia, Netherlands, Pakistan, Poland, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Turkmenistan, United Kingdom of Great Britain and Northern Ireland)
   • Formica lusatica Seifert, 1997
   • Formica rizadaghi Arnoldi, 1968
   • Formica rufibarbis caucasica Wheeler, W.M., 1913

clara sinae Emery, 1925 (China)
   • Formica rufibarbis orientalis Wheeler, W.M., 1923

clarissima Emery, 1925 (China, Mongolia, Russian Federation)
   • Formica subpilosa ruzskyi Dlussky, 1965

coloradensis Creighton, 1940 (United States)
comata Wheeler, W.M., 1909 (United States)
conica Fabricius, 1798
connecticutensis Buckley, 1866
corsica Seifert, 2002 (France)
creightoni Buren, 1968 (United States)
criniventris Wheeler, W.M., 1912 (United States)
   • Formica crinita Wheeler, W.M., 1909

cunicularia Latreille, 1798 (Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Channel Islands, China, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, India, Iran, Italy, Kyrgyzstan, Latvia, Lithuania, Luxembourg, Mongolia, Montenegro, Morocco, Netherlands, Norway, Pakistan, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)
   • Formica cunicularia fuscoides Dlussky, 1967
   • Formica fusca fuscorufibarbis Forel, 1874
   • Formica fusca rubescens Forel, 1904

curiosa Creighton, 1935 (Canada, United States)
   • Formica parcipappa Cole, 1947

D

dachaidanensis Chang & He, 2002 (China)
dakotensis Emery, 1893 (Canada, United States)
   • Formica dakotensis saturata Wheeler, W.M., 1917
   • Formica dakotensis wasmanni Forel, 1904
   • Formica fusca specularis Wheeler, W.M., 1908
   • Formica montigena Wheeler, W.M., 1904

decipiens Bondroit, 1918 (Andorra, France, Iberian Peninsula, Spain)
delinghaensis Chang & He, 2002 (China)
densiventris Viereck, 1903 (Canada, Mexico, United States)
   • Formica microgyna rasilis Wheeler, W.M., 1903
   • Formica rasilis spicata Creighton, 1950

didyma Fabricius, 1782
difficilis Emery, 1893 (United States)
   • Formica difficilis consocians Wheeler, W.M., 1904
   • Formica habrogyna Cole, 1939

dirksi Wing, 1949 (United States)
dislocata Say, 1836
dolosa Buren, 1944 (United States)
dusmeti Emery, 1909 (Iberian Peninsula, Portugal, Spain)

E

elevata Fabricius, 1782
elongata Fabricius, 1787
emeryi Wheeler, W.M., 1913 (Canada, United States)
exsecta Nylander, 1846 (Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, China, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Iberian Peninsula, Italy, Kazakhstan, Kyrgyzstan, Latvia, Liechtenstein, Lithuania, Luxembourg, Mongolia, Netherlands, Norway, Poland, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)
   • Formica dalcqi Bondroit, 1918
   • Formica exsecta etrusca Emery, 1909
   • Formica exsecta exsectopressilabris Forel, 1874
   • Formica exsecta rubens Forel, 1874
   • Formica exsecta sudetica Scholz, 1924
   • Formica kontuniemii Betrem, 1954
   • Formica nemoralis Dlussky, 1964

exsectoides Forel, 1886 (Canada, Mexico, United States)
   • Formica exsectoides davisi Wheeler, W.M., 1913
   • Formica exsectoides hesperia Wheeler, W.M., 1913

exsectorubens Ruzsky, 1905

F

fatale Christ, 1791
fennica Seifert, 2000 (Azerbaijan, Finland, Georgia)
ferocula Wheeler, W.M., 1913 (United States)
foetens Olivier, 1792
   • Formica foetida De Geer De Geer, 1773

foreli Bondroit, 1918 (Albania, Andorra, Austria, Belgium, Croatia, Czech Republic, Denmark, Georgia, Germany, Iberian Peninsula, Norway, Poland, Russian Federation, Spain, Sweden, Switzerland, Turkey)
   • Formica goesswaldi Kutter, 1967
   • Formica naefi Kutter, 1957
   • Formica tamarae Dlussky, 1964

foreliana Wheeler, W.M., 1913 (Mexico, United States)
forsslundi Lohmander, 1949 (Belarus, China, Denmark, Finland, Germany, Mongolia, Norway, Poland, Russian Federation, Sweden, Switzerland, Tibet)
   • Formica brunneonitida Dlussky, 1964
   • Formica forsslundi strawinskii Petal, 1963
   • Formica fossilabris Dlussky, 1965

fossaceps Buren, 1942 (Canada, United States)
francoeuri Bolton, 1995 (Mexico, United States)
   • Formica pilicornis Emery, 1893

frontalis Santschi, 1919 (Andorra, Iberian Peninsula, Kyrgyzstan, Portugal, Spain)
fukaii Wheeler, W.M., 1914 (China, Japan)
fusca Linnaeus, 1758 (Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Canada, Channel Islands, China, Croatia, Denmark, Estonia, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, India, Isle of Man, Latvia, Lithuania, Luxembourg, Malta, Montenegro, Nepal, Netherlands, Norway, Pakistan, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)
   • Formica barbata Razoumowsky, 1789
   • Formica flavipes Geoffroy, 1785
   • Formica fusca pallipes Kuznetsov-Ugamsky, 1926
   • Formica fusca rufipes Stitz, 1930
   • Formica glebaria Nylander, 1846
   • Formica libera Scopoli, 1763
   • Formica tristis Christ, 1791

fusca alpicola Gredler, 1858 (Austria)
fusca fuscolemani Samsinak, 1951 (Czech Republic)
fusca hyrcana Arnoldi, 1968 (Russian Federation)
fusca maura Santschi, 1929 (Morocco)
fusca tombeuri Bondroit, 1917 (Portugal)
fuscescens Gmelin, 1790
fuscicauda Motschoulsky, 1863
fuscocinerea Forel, 1874 (Austria, France, Germany, Hungary, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Switzerland, Ukraine)
   • Formica lefrancoisi Bondroit, 1918

G

gagates Latreille, 1798 (Albania, Austria, Bulgaria, China, Croatia, Czech Republic, France, Germany, Greece, Hungary, Iberian Peninsula, India, Montenegro, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Switzerland, Turkey, Ukraine)
   • Formica capsincola Schilling, 1839
   • Formica fusca fuscogagates Forel, 1874
   • Formica morio Latreille, 1798

gagatoides Ruzsky, 1904 (Belgium, China, Finland, India, Japan, Montenegro, Norway, Russian Federation, Sweden, Switzerland)
georgica Seifert, 2002 (Georgia, Turkey)
gerardi Bondroit, 1917 (Andorra, France, Gibraltar, Iberian Peninsula, Portugal, Spain)
glabra Gmelin, 1790
glabridorsis Santschi, 1925 (China)
glacialis Wheeler, W.M., 1908 (Canada, United States)
   • Formica fusca fairchildi Wheeler, W.M., 1927

glauca Ruzsky, 1896 (Bulgaria, Germany, Iran, Poland, Romania, Russian Federation, Ukraine)
   • Formica rufibarbis katuniensis Ruzsky, 1915

gnava Buckley, 1866 (Mexico, United States)
gravelyi Mukerjee, 1930 (India)
gynocrates Snelling, R.R. & Buren, 1985 (United States)

H

hayashi Terayama & Hashimoto, 1996 (Japan, Republic of Korea)
hewitti Wheeler, W.M., 1917 (Canada, Mexico, United States)
hirta Gravenhorst, 1807

I

impexa Wheeler, W.M., 1905 (Canada, United States)
incerta Buren, 1944 (United States)
incisa Smith, F., 1858
indianensis Cole, 1940 (United States)
inequalis Lowne, 1865
insultans Forskål, 1775
integra Nylander, 1856 (Canada, United States)
   • Formica integra similis Mayr, 1886

integroides Wheeler, W.M., 1913 (Canada, Mexico, United States)
   • Formica rufa subcaviceps Creighton, 1940
   • Formica rufa subfasciata Creighton, 1940

J

japonica Motschoulsky, 1866 (China, Democratic Peoples Republic of Korea, Japan, Mongolia, Republic of Korea, Russian Federation)
   • Formica fusca nipponensis Forel, 1900

K

kashmirica Stärcke, 1935 (India, Kashmir)
knighti Buren, 1944 (United States)
kozlovi Dlussky, 1965 (Mongolia)
kupyanskayae Bolton, 1995 (Democratic Peoples Republic of Korea, Russian Federation)
   • Formica opacus Kupyanskaya, 1980

L

laeviceps Creighton, 1940 (United States)
lasioides Emery, 1893 (Canada, Mexico, United States)
   • Formica lasioides picea Emery, 1895
   • Formica lasioides vetula Wheeler, W.M., 1912

lemani Bondroit, 1917 (Albania, Andorra, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Democratic Peoples Republic of Korea, Finland, France, Georgia, Germany, Greece, Iberian Peninsula, Italy, Japan, Kyrgyzstan, Mongolia, Montenegro, Netherlands, Norway, Poland, Republic of Korea, Republic of Macedonia, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)
   • Formica fusca borealis Vashkevich, 1924

lepida Wheeler, W.M., 1913 (United States)
limata Wheeler, W.M., 1913 (Canada, Mexico, United States)
lincecumii Buckley, 1866
liogaster Chang & He, 2002 (China)
liopthalma Chang & He, 2002 (China)
litoralis Kuznetsov-Ugamsky, 1926 (China, Kazakhstan, Kyrgyzstan)
longipilosa Francoeur, 1973 (United States)
lugubris Zetterstedt, 1838 (Albania, Andorra, Austria, Belarus, Bulgaria, China, Croatia, Czech Republic, Democratic Peoples Republic of Korea, Estonia, Finland, France, Germany, Greece, Iberian Peninsula, Ireland, Italy, Latvia, Mongolia, Montenegro, Norway, Poland, Republic of Korea, Romania, Russian Federation, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)
   • Formica congerens Nylander, 1846
   • Formica pratensis unicolor Ruzsky, 1926
   • Formica rufa alpina Santschi, 1911
   • Formica rufa montana Sadil, 1953
   • Formica rufa nylanderi Bondroit, 1920
   • Formica rufa santschii Wheeler, W.M., 1913

lutea Duméril, 1860

M

maculata Geoffroy, 1785
malabarica Schrank, 1837
maligna Forskål, 1775
manchu Wheeler, W.M., 1929 (China, Mongolia, Russian Federation)
   • Formica dlusskyi Bolton, 1995
   • Formica longiceps Dlussky Dlussky, 1964

manni Wheeler, W.M., 1913 (United States)
maxillosa Fabricius, 1775
melanophthalma Reich, 1793
melanopis Gmelin, 1790
mesasiatica Dlussky, 1964 (China, Russian Federation)
microgyna Wheeler, W.M., 1903 (Canada, Mexico, United States)
   • Formica microgyna recidiva Wheeler, W.M., 1913

microphthalma Francoeur, 1973 (United States)
miniocca Chang & He, 2002 (China)
minuta Lowne, 1865
moki Wheeler, W.M., 1906 (Mexico, United States)
   • Formica rufibarbis occidua Wheeler, W.M., 1913

molestans Latreille, 1802
montana Wheeler, W.M., 1910 (Canada, Mexico, United States)
   • Formica cinerea neocinerea Wheeler, W.M., 1913
   • Formica cinerea rutilans Wheeler, W.M., 1913

montaniformis Kuznetsov-Ugamsky, 1929
montivaga Santschi, 1928
   • Formica montana Kuznetsov-Ugamsky Kuznetsov-Ugamsky, 1923

morsei Wheeler, W.M., 1906 (United States)
mucescens Wheeler, W.M., 1913 (United States)
mutinensis (Canestrini, 1873)

N

nana Latreille, 1802
neoclara Emery, 1893 (Canada, Mexico, United States)
   • Formica fusca pruinosa Wheeler, W.M., 1917

neogagates Viereck, 1903 (Canada, Mexico, United States)
neorufibarbis Emery, 1893 (Canada, Mexico, United States)
   • Formica fusca algida Wheeler, W.M., 1915
   • Formica fusca gelida Wheeler, W.M., 1913

nepticula Wheeler, W.M., 1905 (United States)
nevadensis Wheeler, W.M., 1904 (United States)
nigropratensis Betrem, 1962 (Germany)
nitida Razoumowsky, 1789
nortonii Buckley, 1866
novaanglae Buckley, 1866

O

obscuripes Forel, 1886 (Canada, United States)
   • Formica rufa aggerans Wheeler, W.M., 1912
   • Formica rufa melanotica Creighton, 1940

obscuriventris Mayr, 1870 (Canada, United States)
   • Formica dryas Wheeler, W.M., 1905
   • Formica dryas gymnomma Wheeler, W.M., 1905
   • Formica rufa clivia Creighton, 1940

obsidiana Emery, 1923 (Taiwan)
obsoleta Linnaeus, 1758
obtusopilosa Emery, 1893 (Canada, Mexico, United States)
   • Formica munda Wheeler, W.M., 1905
   • Formica munda alticola Wheeler, W.M., 1917

occidentalis Buckley, 1866
occulta Francoeur, 1973 (Mexico, United States)
omnivora Linnaeus, 1758
opaciventris Emery, 1893 (Canada, United States)
orangea Seifert & Schultz, 2009 (Afghanistan, Kyrgyzstan, Uzbekistan)
oreas Wheeler, W.M., 1903 (Canada, United States)
   • Formica oreas comptula Wheeler, W.M., 1913

oregonensis Cole, 1938 (United States)
ovata Reich, 1793

P

pachucana Francoeur, 1973 (Mexico)
pacifica Francoeur, 1973 (United States)
pallida (Canestrini, 1873)
pallidefulva Latreille, 1802 (Canada, United States)
   • Formica pallidefulva delicata Cole, 1938
   • Formica pallidefulva fuscata Emery, 1893
   • Formica pallidefulva nitidiventris Emery, 1893
   • Formica pallidefulva succinea Wheeler, W.M., 1904
   • Formica schaufussi Mayr, 1866

pallidelutea Latreille, 1802
pallipes Latreille, 1787
pamirica Dlussky, 1965 (China, Kyrgyzstan, Pakistan, Turkmenistan)
paralugubris Seifert, 1996 (Albania, Andorra, Austria, Canada, Germany, Iberian Peninsula, Spain, Switzerland)
pergandei Emery, 1893 (United States)
   • Formica sanguinea sublucida Creighton, 1950

perpilosa Wheeler, W.M., 1913 (Canada, Mexico, United States)
persica Seifert & Schultz, 2009 (Iran)
phyllophila Jerdon, 1851
picea Nylander, 1846 (Andorra, Belgium, Bulgaria, China, Czech Republic, Finland, Georgia, Greece, Iberian Peninsula, India, Italy, Kazakhstan, Pakistan, Poland, Russian Federation, Spain, Turkey, United Kingdom of Great Britain and Northern Ireland)
   • Formica fusca inplana Emery, 1925
   • Formica fusca orientalis Ruzsky, 1915
   • Formica fusca piceoinplana Emery, 1925
   • Formica glabra White, W.F. White, W.F., 1884
   • Formica picea lochmatteri Stärcke, 1935

picipes Reich, 1793
pisarskii Dlussky, 1964 (Mongolia)
planipilis Creighton, 1940 (Canada, United States)
podzolica Francoeur, 1973 (Canada, Mexico, United States)
politurata Buckley, 1866
polyctena Foerster, 1850 (Albania, Andorra, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Germany, Greece, Hungary, Iberian Peninsula, India, Italy, Kazakhstan, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Republic of Moldova, Romania, Russian Federation, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Ukraine)
   • Formica major Nylander, 1849
   • Formica rufa minor Gösswald, 1951
   • Formica rufa nuda Karavaiev, 1930

postoculata Kennedy & Dennis, 1937 (United States)
pratensis Retzius, 1783 (Albania, Andorra, Austria, Belarus, Belgium, Bulgaria, Channel Islands, China, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Georgia, Germany, Greece, Guernsey, Hungary, Iberian Peninsula, Iran, Italy, Kazakhstan, Kyrgyzstan, Latvia, Lithuania, Luxembourg, Mongolia, Montenegro, Netherlands, Norway, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)
   • Formica minor pratensoides Gösswald, 1951
   • Formica pratensis cordieri Bondroit, 1917
   • Formica rufa ciliata Ruzsky, 1926
   • Formica rufa grouvellei Bondroit, 1918
   • Formica rufa nigricans Bondroit, 1912

pratensis nuda Ruzsky, 1926 (Russian Federation)
pratensis rufoides Stärcke, 1926 (Switzerland)
pratensis staerckei Betrem, 1960 (Netherlands)
pressilabris Nylander, 1846 (Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, Czech Republic, Denmark, Estonia, Finland, France, Germany, Hungary, Iberian Peninsula, Latvia, Lithuania, Luxembourg, Mongolia, Netherlands, Norway, Poland, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine)
   • Formica exsecta rufomaculata Ruzsky, 1895

prociliata Kennedy & Dennis, 1937 (United States)
propatula Francoeur, 1973 (Mexico)
propinqua Creighton, 1940 (United States)
puberula Emery, 1893 (United States)
pulla Francoeur, 1973 (Mexico)
pusilla De Geer, 1773
pyrenaea Bondroit, 1918 (France)

Q

quadrinotata Losana, 1834
querquetulana Kennedy & Dennis, 1937 (United States)

R

ravida Creighton, 1940 (Canada, United States)
   • Formica rufa haemorrhoidalis Creighton, 1940
   • Formica rufa tahoensis Creighton, 1940

rediana Leach, 1825
reflexa Buren, 1942 (Canada, United States)
retecta Francoeur, 1973 (Honduras, Mexico)
robusta Chang & He Chang & He, 2002 (China)
rostrata Fabricius, 1787
rubescens Leach, 1825
rubicunda Emery, 1893 (Canada, Mexico, United States)
rufa Linnaeus, 1761 (Albania, Andorra, Armenia, Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, Italy, Kazakhstan, Latvia, Lithuania, Luxembourg, Malta, Montenegro, Netherlands, Norway, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Serbia, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland, Uzbekistan)
   • Formica apicalis Smith, F., 1858
   • Formica dorsata Panzer, 1798
   • Formica ferruginea Christ Christ, 1791
   • Formica gaullei Bondroit, 1917
   • Formica piniphila Schenck, 1852
   • Formica rufa meridionalis Nasonov, 1889
   • Formica rufa rufopratensis Forel, 1874

rufa angusticeps Stärcke, 1947 (Netherlands)
rufa brevisetosa Ruzsky, 1926 (Russian Federation, Ukraine)
rufa constricta Karavaiev, 1929 (Russian Federation)
rufa obscurata Santschi, 1925 (France)
rufa rufopratensoides Forel, 1874 (Switzerland)
rufa tshugunovi Ruzsky, 1914 (Russian Federation)
rufibarbis Fabricius, 1793 (Albania, Andorra, Armenia, Austria, Balearic Islands, Belarus, Belgium, Bosnia and Herzegovina, Bulgaria, China, Croatia, Czech Republic, Denmark, Estonia, Finland, France, Georgia, Germany, Greece, Hungary, Iberian Peninsula, India, Iran, Italy, Latvia, Lithuania, Luxembourg, Montenegro, Netherlands, Norway, Poland, Portugal, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Turkmenistan, Ukraine, United Kingdom of Great Britain and Northern Ireland)
   • Formica defensor Smith, F., 1878
   • Formica fraterna Smith, F., 1878
   • Formica fusca cinereorufibarbis Forel, 1874
   • Formica nicaeensis Leach, 1825
   • Formica rufibarbis piligera Lomnicki, 1925

rufibarbis clarorufibarbis Wheeler, W.M. & Mann, 1916 (Russian Federation)
rufibarbis subpilosorufibarbis Ruzsky, 1905 (Russian Federation)
ruficeps Motschoulsky, 1863 (Indonesia)
ruficornis Gmelin, 1790
rufolucida Collingwood, 1962 (Myanmar)
rupestris Leach, 1825

S

saccharivora Linnaeus, 1758
sanguinea Latreille, 1798 (Afghanistan, Albania, Andorra, Armenia, Austria, Balearic Islands, Belarus, Belgium, Bulgaria, Canary Islands, China, Croatia, Czech Republic, Democratic Peoples Republic of Korea, Denmark, Estonia, Finland, France, Georgia, Germany, Gibraltar, Greece, Hungary, Iberian Peninsula, India, Iran, Japan, Kazakhstan, Kyrgyzstan, Latvia, Lithuania, Luxembourg, Mongolia, Netherlands, Norway, Pakistan, Poland, Portugal, Republic of Korea, Republic of Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Turkey, Ukraine, United Kingdom of Great Britain and Northern Ireland)
   • Formica dominula Nylander, 1846
   • Formica sanguinea arenicola Kuznetsov-Ugamsky, 1928
   • Formica sanguinea borea Santschi, 1925
   • Formica sanguinea clara Karavaiev, 1927
   • Formica sanguinea clarior Ruzsky, 1905
   • Formica sanguinea flavorubra Forel, 1909
   • Formica sanguinea fusciceps Emery, 1895
   • Formica sanguinea griseopubescens Kuznetsov-Ugamsky, 1926
   • Formica sanguinea leninei Santschi, 1928
   • Formica sanguinea mollesonae Ruzsky, 1903
   • Formica sanguinea monticola Kuznetsov-Ugamsky, 1926
   • Formica sanguinea rotundata Kuznetsov-Ugamsky, 1926
   • Formica sanguinea strennua Santschi, 1925
   • Formica sanguinea tristis Karavaiev, 1929

saxicola Buckley, 1866
schardj Forskål, 1775
scitula Wheeler, W.M., 1913 (United States)
selysi Bondroit, 1918 (Albania, Austria, France, Germany, Iberian Peninsula, Slovenia, Spain, Switzerland)
   • Formica torrentium Bernard, 1967

sentschuensis Ruzsky, 1915 (China, Tibet)
   • Formica dalailamae Ruzsky, 1915

siamicarubra Christ, 1791
sibylla Wheeler, W.M., 1913 (United States)
sinensis Wheeler, W.M., 1913 (China)
spatulata Buren, 1944 (United States)
stenoptera Foerster, 1850
subaenescens Emery, 1893 (Canada, Mexico, United States)
   • Formica fusca marcida Wheeler, W.M., 1913

subcyanea Wheeler, W.M., 1913 (Mexico)
subelongata Francoeur, 1973 (United States)
subintegra Wheeler, W.M., 1908 (Canada, United States)
subnitens Creighton, 1940 (Canada, United States)
subpicea Motschoulsky, 1863
subpilosa Ruzsky, 1902 (Afghanistan, Armenia, Azerbaijan, China, Georgia, Kyrgyzstan, Russian Federation, Turkey, Turkmenistan, Uzbekistan)
   • Formica cinerea bipilosa Karavaiev, 1926

subpolita Mayr, 1886 (Canada, Mexico, United States)
   • Formica flammiventris Wheeler, W.M., 1912
   • Formica rufiventris Emery Emery, 1893
   • Formica subpolita camponoticeps Wheeler, W.M., 1913
   • Formica subpolita ficticia Wheeler, W.M., 1913

subsericea Say, 1836 (United States)
   • Formica lecontei Kennedy & Dennis, 1937

subspinosa Buckley, 1866
suecica Adlerz, 1902 (Austria, Finland, Norway, Russian Federation, Sweden)

T

talbotae Wilson, 1977 (United States)
tarimica Seifert & Schultz, 2009 (China)
tenuissima Buckley, 1866
testacea Gmelin, 1790
testaceipes Leach, 1825
thoracica Olivier, 1792
tianshanica Seifert & Schultz, 2009 (China, Kyrgyzstan)
tomentosa Reich, 1793
transkaucasica Nasonov, 1889 (Denmark)
   • Formica gagates piceogagates Karavaiev, 1926

transmontanis Francoeur, 1973 (Canada, United States)
triangularis Say, 1836
truncorum Fabricius, 1804 (Austria, Belarus, Belgium, Bulgaria, China, Croatia, Czech Republic, Democratic Peoples Republic of Korea, Denmark, Estonia, Finland, Germany, Hungary, India, Japan, Kazakhstan, Kyrgyzstan, Latvia, Lithuania, Mongolia, Netherlands, Norway, Pakistan, Poland, Republic of Korea, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Sweden, Switzerland, Turkey, Ukraine)
   • Formica rufa menozzii Stitz, 1939
   • Formica rufa rufotruncicola Ruzsky, 1896
   • Formica rufa staegeri Stitz, 1939
   • Formica rufa truncicolopratensis Forel, 1874
   • Formica simulata Smith, F., 1878
   • Formica truncicola Nylander, 1846

truncorum finzii Stitz, 1939 (Germany)

U

ulkei Emery, 1893 (Canada, United States)
   • Formica ulkei hebescens Wheeler, W.M., 1913

uralensis Ruzsky, 1895 (Belarus, China, Democratic Peoples Republic of Korea, Denmark, Estonia, Finland, France, Germany, Kazakhstan, Latvia, Lithuania, Mongolia, Norway, Poland, Russian Federation, Sweden, Switzerland, Ukraine)
   • Formica pratensis superba Wheeler, W.M., 1933

V

vagans Fabricius, 1793
venosa Gmelin, 1790
villiscapa Chang & He, 2002 (China)
vinculans Wheeler, W.M., 1913 (United States)
virginiana Buckley, 1866
volgensis Ruzsky, 1914

W

wheeleri Creighton Creighton, 1935 (Mexico, United States)
wheeleri Stitz Stitz, 1939
wongi Wu, J., 1990 (China)

X

xerophila Smith, M.R., 1939 (Mexico, United States)
   • Formica moki grundmanni Cole, 1943

Y

yessensis Wheeler, W.M., 1913 (China, Democratic Peoples Republic of Korea, Japan, Republic of Korea, Russian Federation)
yoshiokae Wheeler, W.M., 1933 (Japan)

Fossil Species

A

aemula Heer, 1867 (Burdigalian, Early Miocene

Radoboj, Croatia)
alsatica Théobald, 1937 (Early Oligocene

Kleinkems, Germany)
annosa LaPolla & Greenwlalt, 2015 (Lutetian, Middle Eocene

Kishenehn Formation shale, Montana, United States)
arcana Scudder, 1877 (Early Miocene?

Quesnel, British Columbia, Canada)
auxillacensis Piton, 1935 (Early Oligocene

Auxillac, Auvergne, France)

B

bauckhorni Meunier, 1915 (Late Oligocene

Rott, Westphalia, Germany)
biamoensis Dlussky, Rasnitsyn & Perfilieva, 2015 (Priabonian, Late Eocene

Bol’shaya Svetlovodnaya, Sikhote-Alin, Russia)
buphthalma Novak, O., 1878 (Late Burdigalian, Early Miocene

Mokrina (Krottensee), Czech Republic)

C

cantalica Piton, 1935 (Late Miocene

Joursac, Auvergne, France)
ceps Zhang, J., 1989 (Early Miocene

Shanwang, China)
cockerelli Carpenter, 1930 (Late Eocene

Florissant, Colorado, United States)

D

demersa Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)

E

eoptera Cockerell, 1923 (Late Eocene

Mossy Creek, near Wellborn, Texas, United States)

F

flavifemoralis Zhang, J., Sun & Zhang, X., 1994 (Early Miocene

Shanwang, China)
flori Mayr, 1868 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)
fragilis Heer, 1867 (Burdigalian, Early Miocene

Radoboj, Croatia)

G

gibbosa Presl, 1822 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)
globiventris Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
grandis Carpenter, 1930 (Late Eocene

Florissant, Colorado, United States)
gravida Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
gustawi Dlussky, 2002 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)

H

heteroptera Cockerell, 1920 (Bartonian, Middle Eocene

Bournemouth, Dorset, U.K.)
horrida Wheeler, W.M., 1915 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)

I

immersa Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)

K

kollari Heer, 1867 (Burdigalian, Early Miocene

Radoboj, Croatia)
kutscheri Dlussky, 2008 (Bartonian, Middle to Late Eocene

Bitterfeld amber, Sachsen-Anhalt, Germany)

L

latinodosa Théobald, 1937 (Chattian stage, Late Oligocene

Aix-en-Provence, France)
lavateri Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
lavateri major Heer, 1867 (Burdigalian, Early Miocene

Radoboj, Croatia)
linquensis Zhang, J., 1989 (Early Miocene

Shanwang, China)
longicollis Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
lucida Giebel, 1856 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)
luteola Presl, 1822 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)

M

macrognatha Presl, 1822 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)
macrophthalma Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
maculipennis Théobald, 1935 (Early Oligocene

Auxillac, Auvergne, France)
martynovi (Popov, 1933) (Serravallian, Miocene

Ormety, Georgia)

N

nigra Presl, 1822 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)

O

oblita Heer, 1867 (Burdigalian, Early Miocene

Radoboj, Croatia)
obscura Heer, 1849 (Burdigalian, Early Miocene

Radoboj, Croatia)
obtecta Heer, 1849 (Burdigalian, Early Miocene

Radoboj, Croatia)
obvoluta Heer, 1849 (Burdigalian, Early Miocene

Radoboj, Croatia)
oculata Heer, 1849 (Burdigalian, Early Miocene

Radoboj, Croatia)
orbata Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
ovala Zhang, J., 1989 (Early Miocene

Shanwang, China)

P

palaeopolonica Dlussky, 2008 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)
paleosibirica Dlussky, Rasnitsyn & Perfilieva, 2015 (Priabonian, Late Eocene

Bol’shaya Svetlovodnaya, Sikhote-Alin, Russia)
parexsecta Dlussky & Putyatina, 2014 (Burdigalian, Early Miocene

Radoboj, Croatia)
parvula Presl, 1822 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)
phaethusa Wheeler, W.M., 1915 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)
pitoni Théobald, 1935 (Pliocene

Lac Chambon, Auvergne, France)
primitiva Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
primordialis Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
procera Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
pulchella Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)

Q

quadrata Holl, 1829 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)

R

radchenkoi Dlussky, 2008 (Priabonian, Late Eocene

Rovno amber, Ukraine)
robusta Carpenter Carpenter, 1930 (Late Eocene

Florissant, Colorado, United States)

S

sepulta Théobald, 1937 (Chattian stage, Late Oligocene

Aix-en-Provence, France)
serresi Théobald, 1937 (Chattian stage, Late Oligocene

Aix-en-Provence, France)
seuberti Heer, 1849 (Messinian, Late Miocene

Oeningen, Switzerland)
strangulata Wheeler, W.M., 1915 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)
surinamensis Berendt, 1830 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)

T

trigona Presl, 1822 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)
tripartita Théobald, 1937 (Early Oligocene

Kleinkems, Germany)

U

ungeri Heer, 1849 (Burdigalian, Early Miocene

Radoboj, Croatia)

ungeri neutra Heer, 1849 (Fossil (details uncertain))

Z

zherikhini Dlussky, 2008 (Bartonian, Middle to Late Eocene

Baltic amber, Baltic Sea region)