Checklist of Pseudomyrmex species

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Pseudomyrmex
Pseudomyrmecinae
Pseudomyrmex gracilis casent0103874 head 1.jpg
Pseudomyrmex gracilis
Pseudomyrmex Distribution.png
Distribution
Pseudomyrmex Species Richness.png
Species Richness
Statistics
Valid Species 155
Invalid Species 139
Fossils 13
Countries Occupied 37
Year Described 1831
First Species Described 1804
Last Species Described 2022

The following species and subspecies belong to the genus Pseudomyrmex. Synonyms are listed under their senior names. For valid names only see Pseudomyrmex species and for distribution information see Pseudomyrmex species by Country.

A

acanthobius (Emery, 1896) (Argentina, Bolivia, Brazil, Paraguay)
   • Pseudomyrma acanthobia fuscata Emery, 1896

acanthobius cocae (Santschi, 1922) (Argentina)
acanthobius virgo (Santschi, 1922) (Argentina, Brazil)
adustus (Borgmeier, 1929) (Brazil)
alternans (Santschi, 1936) (French Guiana)
alustratus Ward, 1989 (Ecuador, Peru)
alvarengai Kempf, 1961 (Brazil)
antiguanus (Enzmann, E.V., 1944) (Guatemala)
antiguanus brunnipes (Enzmann, E.V., 1944) (Guatemala)
apache Creighton, 1953 (Mexico, United States)
arcanus Ward & Branstetter, 2022 (Mexico)
atripes (Smith, F., 1860) (Bolivia, Brazil, Colombia, Ecuador)
   • Pseudomyrma levigata kitschelti Forel, 1911

B

beccarii (Menozzi, 1935) (Costa Rica, Guyana)
boopis (Roger, 1863) (Colombia, Costa Rica, Ecuador, Guatemala, Honduras, Mexico, Panama, Trinidad and Tobago, Venezuela)
   • Pseudomyrma excavata Mayr, 1870
   • Pseudomyrma excavata flaviventris Emery, 1896
   • Pseudomyrma excavata fusciceps Santschi, 1931
   • Pseudomyrma modesta Smith, F., 1862
   • Pseudomyrma tenuis guatemalensis Enzmann, E.V., 1944
   • Pseudomyrma thoracica Norton, 1868

browni Kempf, 1967 (Brazil, Colombia, Panama)
brunneus (Smith, F., 1877) (Costa Rica, Greater Antilles, Mexico)
   • Pseudomyrma brunnea nigrita Enzmann, E.V., 1944

C

caeciliae (Forel, 1913) (Costa Rica, Greater Antilles, Guatemala, Honduras, Mexico, Nicaragua, Panama, United States)
capillatus Ward & Branstetter, 2022 (Mexico)
championi (Forel, 1899) (Mexico)
   • Pseudomyrma leonhardi Stitz, 1937

cladoicus (Smith, F., 1858) (Brazil)
cognatus Ward & Branstetter, 2022 (El Salvador, Guatemala, Honduras, Mexico, Nicaragua)
colei (Enzmann, E.V., 1944) (Ecuador, Peru)
colei vistanus (Enzmann, E.V., 1944) (Peru)
comitator Ward & Branstetter, 2022 (Mexico)
concolor (Smith, F., 1860) (Brazil, French Guiana, Guyana, Suriname)
   • Pseudomyrma damnosa Wheeler, W.M., 1921
   • Pseudomyrma latinoda Mayr, 1878
   • Pseudomyrma latinoda endophyta Forel, 1912

cordiae (Forel, 1904) (Bolivia, Peru)
cretus Ward, 1989 (Costa Rica, Guatemala)
crudelis Ward, 1999 (Peru)
cubaensis (Forel, 1901) (Argentina, Bahamas, Bolivia, Brazil, Cayman Islands, Colombia, Costa Rica, Cuba, Dominican Republic, Greater Antilles, Guatemala, Guyana, Haiti, Jamaica, Mexico, Panama, Paraguay, Peru, United States, Venezuela)
curacaensis (Forel, 1912) (Bolivia, Colombia, French Guiana, Guadeloupe, Guyana, Lesser Antilles, Panama, Suriname, Trinidad and Tobago, Venezuela)

D

deminutus Ward, 1999 (Brazil)
dendroicus (Forel, 1904) (Brazil, Colombia, Ecuador, Peru)
   • Pseudomyrma dendroica emarginata Forel, 1904

denticollis (Emery, 1890) (Argentina, Brazil, Paraguay)
depressus (Forel, 1906) (Colombia)
distinctus (Smith, F., 1877) (Mexico)
   • Pseudomyrma distincta pulchella Forel, 1899

duckei (Forel, 1906) (Brazil, Costa Rica, Ecuador, Guatemala)

E

eculeus Ward, 1999 (Ecuador, Peru)
eduardi (Forel, 1912) (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Greater Antilles, Jamaica, Panama, Paraguay, Venezuela)
ejectus (Smith, F., 1858) (Bolivia, Brazil, Costa Rica, Greater Antilles, Guatemala, Mexico, United States)
   • Ponera lincecumii Buckley, 1866

elongatulus (Dalla Torre, 1892) (Belize, Guatemala, Mexico, United States)
   • Pseudomyrma decipiens Forel, 1899
   • Pseudomyrma elongata Smith, F., 1877

elongatus (Mayr, 1870) (Belize, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Ecuador, El Salvador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, United States, Venezuela)
   • Pseudomyrma allidora Enzmann, E.V., 1944
   • Pseudomyrma alliodorae Wheeler, W.M., 1942
   • Pseudomyrma elongata tandem Forel, 1906

ereptor Ward & Branstetter, 2022 (Mexico)
ethicus (Forel, 1911) (Bolivia, Brazil, Ecuador)
euryblemma (Forel, 1899) (Colombia, Costa Rica, Guatemala, Mexico, Paraguay)
   • Pseudomyrma kurokii Forel, 1906

evitus Ward, 2017 (Belize, Costa Rica, Guatemala, Honduras, Mexico)
excisus (Mayr, 1870) (Colombia)
exoratus Ward & Branstetter, 2022 (Honduras, Mexico)

F

faber (Smith, F., 1858) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Panama, Suriname)
   • Pseudomyrma perforator Smith, F., 1860

fasciatus Ward & Branstetter, 2022 (Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua)
feralis Ward, 2017 (Guatemala)
ferox Ward, 1999 (Brazil, Colombia, Peru)
ferrugineus (Smith, F., 1877) (Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua)
   • Pseudomyrma belti bequaerti Enzmann Enzmann, E.V., 1944
   • Pseudomyrma belti bequaerti Wheeler, W.M. Wheeler, W.M., 1942
   • Pseudomyrma belti fulvescens Emery, 1890
   • Pseudomyrma belti saffordi Wheeler, W.M., 1942
   • Pseudomyrma belti vesana Wheeler, W.M., 1942
   • Pseudomyrma canescens Wasmann Wasmann, 1915
   • Pseudomyrma kuenckeli hondurana Enzmann, E.V., 1944
   • Pseudomyrma wasmanni Wheeler, W.M., 1921

fervidus (Smith, F., 1877) (Mexico)
fiebrigi (Forel, 1908) (Argentina, Paraguay)
filiformis (Fabricius, 1804) (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Mexico, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)
   • Pseudomyrma biconvexa Forel, 1899
   • Pseudomyrma biconvexa longiceps Forel, 1906
   • Pseudomyrma cephalica Smith, F., 1855
   • Pseudomyrma longiceps Stitz, 1933

flavicornis (Smith, F., 1877) (Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua)
   • Pseudomyrma belti Emery, 1890
   • Pseudomyrma belti fellosa Wheeler, W.M., 1942
   • Pseudomyrma belti obnubila Menozzi, 1927

flavidulus (Smith, F., 1858) (Argentina, Bahamas, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, Greater Antilles, Grenada, Guadeloupe, Guatemala, Lesser Antilles, Mexico, Panama, Paraguay, Trinidad and Tobago, Venezuela)
fortis (Forel, 1899) (Colombia, Costa Rica, Guatemala, Mexico, Panama, Venezuela)

G

gebellii (Forel, 1899) (Colombia, Panama)
gibbinotus (Forel, 1908) (Brazil)
godmani (Forel, 1899) (Brazil, Colombia, Costa Rica, Panama)
   • Pseudomyrmex semotus Kempf, 1960

goeldii (Forel, 1912) (Brazil)
gracilis (Fabricius, 1804) (Argentina, Barbados, Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, El Salvador, French Guiana, Greater Antilles, Guadeloupe, Guatemala, Guyana, Hawaii, Honduras, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, United States, Uruguay, Venezuela)
   • Pseudomyrma bicolor Guérin-Méneville, 1844
   • Pseudomyrma canescens Smith, F. Smith, F., 1877
   • Pseudomyrma dimidiata Roger, 1863
   • Pseudomyrma gracilis atrinoda Santschi, 1934
   • Pseudomyrma gracilis glabriventris Santschi, 1922
   • Pseudomyrma gracilis longinoda Enzmann, E.V., 1944
   • Pseudomyrma gracilis velifera Stitz, 1933
   • Pseudomyrma mexicana Roger, 1863
   • Pseudomyrma pilosula Smith, F., 1877
   • Pseudomyrma sericata Smith, F., 1855
   • Pseudomyrma variabilis Smith, F., 1877
   • Pseudomyrma volatilis Smith, F., 1877

gracilis argentinus (Santschi, 1934) (Argentina)

H

haytianus (Forel, 1901) (Dominican Republic, Greater Antilles, Haiti)
   • Pseudomyrma haytiana affinis Wheeler, W.M., 1936
   • Pseudomyrma haytiana torquata Wheeler, W.M., 1936

hesperius Ward, 1993 (Mexico)
holmgreni (Wheeler, W.M., 1925) (Paraguay, Peru, Trinidad and Tobago)
hospitalis Ward, 1999 (Peru)

I

incurrens (Forel, 1912) (Brazil)
inquilinus Ward, 1996 (Argentina)
insuavis Ward, 1999 (Colombia)
ita (Forel, 1906) (Belize, Brazil, Colombia, Costa Rica, El Salvador, Guatemala, Honduras, Mexico, Nicaragua, Panama, Paraguay)
   • Pseudomyrma sericea acaciarum Wheeler, W.M., 1942
   • Pseudomyrma sericea acaciorum Enzmann, E.V., 1944

J

janzeni Ward, 1993 (Mexico)

K

kuenckeli (Emery, 1890) (Argentina, Bolivia, Brazil, Colombia, Costa Rica, Honduras, Mexico, Panama, Paraguay, Peru, Trinidad and Tobago)
   • Pseudomyrma crenulata Enzmann, E.V., 1944
   • Pseudomyrma kuenckeli bierigi Santschi, 1932
   • Pseudomyrma kuenckeli dichroa Forel, 1904

L

laevifrons Ward, 1989 (Brazil, Costa Rica, Ecuador)
   • Pseudomyrma laeviceps Smith, F., 1877

laevigatus (Smith, F., 1877) (Brazil, Colombia, Costa Rica, French Guiana, Guyana, Suriname)
laevivertex (Forel, 1906) (Brazil, Colombia, Paraguay, Trinidad and Tobago)
leptosus Ward, 1985 (United States)
lisus (Enzmann, E.V., 1944) (Colombia, Costa Rica, Honduras, Paraguay, Peru)
lizeri (Santschi, 1922) (Argentina, Bolivia, Paraguay)
longior (Forel, 1904) (Brazil, Peru)
longus (Forel, 1912) (Brazil, Paraguay)
lynceus (Spinola, 1851) (Chile)

M

maculatus (Smith, F., 1855) (Argentina, Barbados, Brazil, Guyana, Peru, Suriname, Trinidad and Tobago)
   • Pseudomyrma nigropilosa laticeps Forel, 1906
   • Pseudomyrma nigropilosa wagneri Santschi, 1922

major (Forel, 1899) (Mexico)
malignus (Wheeler, W.M., 1921) (French Guiana, Guyana, Suriname, Trinidad and Tobago)
   • Pseudomyrma auripes Wheeler, W.M., 1922
   • Pseudomyrma maligna cholerica Wheeler, W.M., 1921
   • Pseudomyrma maligna crucians Wheeler, W.M., 1921

mandibularis (Spinola, 1851) (Brazil)
micans Ward, 2017 (Bolivia, Peru)
mixtecus Ward, 1993 (Mexico)
monochrous (Dalla Torre, 1892) (Brazil)
   • Pseudomyrma unicolor Smith, F., 1877

mordax (Warming, 1894) (Colombia, Panama, Venezuela)
   • Pseudomyrma arborissanctae symbiotica Forel, 1904

N

niger (Donisthorpe, 1940) (Bolivia, Guyana)
nigrocinctus (Emery, 1890) (Costa Rica, El Salvador, French Guiana, Guatemala, Guyana, Nicaragua, Suriname)
   • Pseudomyrma alfari Forel, 1906
   • Pseudomyrma nigrocincta bicincta Santschi, 1922
   • Pseudomyrma peltata Menozzi, 1927

nigropilosus (Emery, 1890) (Costa Rica, Mexico)
nimbus Ward & Branstetter, 2022 (Costa Rica)

O

obtusus Ward, 2017 (Brazil, Costa Rica, Panama, Peru, Suriname, Venezuela)
oculatus (Smith, F., 1855) (Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guatemala, Guyana, Honduras, Mexico, Nicaragua, Panama, Peru, Suriname, Trinidad and Tobago, Venezuela)
   • Pseudomyrma advena Smith, F., 1855
   • Pseudomyrma dolichopsis Forel, 1899
   • Pseudomyrma dolichopsis implicata Forel, 1911
   • Pseudomyrma sericea altinoda Mann, 1916
   • Pseudomyrma wessoni Enzmann, E.V., 1944
   • Pseudomyrma wessoni tuberculata Enzmann, E.V., 1944

oki (Forel, 1906) (Colombia, Costa Rica, Panama)
opaciceps Ward, 1993 (El Salvador, Guatemala, Mexico)
osurus (Forel, 1911) (Costa Rica, Panama)
   • Pseudomyrma laevigata insularis Enzmann, E.V., 1944

P

pallens (Mayr, 1870) (Colombia, Costa Rica, Mexico, Peru, Venezuela)
pallens landolti (Forel, 1912) (Colombia)
pallidus (Smith, F., 1855) (Argentina, Belize, Bolivia, Brazil, Colombia, Costa Rica, Cuba, Greater Antilles, Guatemala, Honduras, Mexico, Paraguay, Trinidad and Tobago, United States, Uruguay)
particeps Ward, 1993 (Costa Rica)
parvulus Ward, 2017 (Bolivia, Brazil)
pazosi (Santschi, 1909) (Cuba, Greater Antilles)
   • Pseudomyrma flavidula jaumei Aguayo, 1932

penetrator (Smith, F., 1877) (Brazil, Ecuador, Guyana, Suriname, Venezuela)
   • Pseudomyrma latinoda coronata Enzmann Enzmann, E.V., 1944
   • Pseudomyrma latinoda coronata Wheeler, W.M. Wheeler, W.M., 1942
   • Pseudomyrma latinoda nigrescens Forel, 1904

peperi (Forel, 1913) (Belize, El Salvador, Guatemala, Honduras, Mexico, Nicaragua)
   • Pseudomyrma sabanica saffordi Enzmann, E.V., 1944
   • Pseudomyrma spinicola convarians Forel, 1913

perboscii (Guérin-Méneville, 1844) (Brazil, Costa Rica, Mexico, Peru, Trinidad and Tobago)
   • Pseudomyrma icterica Wheeler, W.M., 1922
   • Pseudomyrma simoides Forel, 1911
   • Tetraponera testacea Smith, F., 1852

peruvianus (Wheeler, W.M., 1925) (Ecuador, Peru)
phyllophilus (Smith, F., 1858) (Argentina, Brazil, Paraguay, Trinidad and Tobago, Uruguay)
   • Pseudomyrma championi paulina Wheeler, W.M., 1916
   • Pseudomyrma mutica Mayr, 1887

pictus (Stitz, 1913) (Bolivia, Brazil, Colombia, Peru)
   • Pseudomyrma picta casta Wheeler, W.M., 1942
   • Pseudomyrma picta heterogyna Wheeler, W.M. & Mann, 1942
   • Pseudomyrma picta humboldi Enzmann, E.V., 1944

pisinnus Ward, 1989 (Brazil, Peru)
pupa (Forel, 1911) (Bolivia, Brazil, Ecuador, Mexico)

R

reconditus Ward, 1993 (Nicaragua)
rochai (Forel, 1912) (Brazil, Colombia, Ecuador, Panama)
rubiginosus (Stitz, 1913) (Brazil, Colombia, Peru)
   • Pseudomyrma sericea huberi Santschi, 1922

rufiventris (Forel, 1911) (Paraguay)
rufomedius (Smith, F., 1877) (Guatemala, Mexico)
   • Pseudomyrma stolli Forel, 1912

S

salvini (Forel, 1899) (Belize, Guatemala, Honduras, Mexico)
santschii (Enzmann, E.V., 1944) (Peru)
satanicus (Wheeler, W.M., 1942) (Costa Rica, Panama)
schuppi (Forel, 1901) (Brazil, Paraguay)
   • Pseudomyrma schuppi confusior Forel, 1901
   • Pseudomyrma schuppi geraensis Forel, 1912

seminole Ward, 1985 (Cuba, Mexico, United States)
sericeus (Mayr, 1870) (Belize, Brazil, Colombia, Costa Rica, Ecuador, Guatemala, Mexico, Panama, Paraguay, Trinidad and Tobago)
simplex (Smith, F., 1877) (Argentina, Bahamas, Belize, Brazil, Colombia, Costa Rica, Cuba, Dominican Republic, Ecuador, El Salvador, Greater Antilles, Grenada, Guatemala, Haiti, Honduras, Jamaica, Mexico, Panama, Paraguay, Peru, Puerto Rico, Trinidad and Tobago, United States, Venezuela)
   • Pseudomyrma delicatula Forel, 1899
   • Pseudomyrma delicatula capperi Forel, 1899
   • Pseudomyrma delicatula panamensis Forel, 1899

simulans Kempf, 1958 (Panama)
solisi (Santschi, 1916) (Argentina)
solisi belgranoi (Santschi, 1916) (Argentina)
spiculus Ward, 1989 (Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Mexico, Panama, Peru, Suriname)
spinicola (Emery, 1890) (Belize, Colombia, Costa Rica, El Salvador, Honduras, Mexico, Nicaragua, Panama)
   • Pseudomyrma spinicola atrox Forel, 1912
   • Pseudomyrma spinicola gaigei Forel, 1914
   • Pseudomyrma spinicola infernalis Wheeler, W.M., 1942
   • Pseudomyrma spinicola scelerosa Wheeler, W.M., 1942
   • Pseudomyrma spinolae infernalis Enzmann, E.V., 1944
   • Pseudomyrma spinolae scelerosa Enzmann, E.V., 1944

squamifer (Emery, 1890) (Brazil)
subater (Wheeler, W.M. & Mann, 1914) (Cuba, Dominican Republic, Ecuador, Greater Antilles, Guatemala, Haiti, Mexico, Paraguay, Trinidad and Tobago)
subtilissimus (Emery, 1890) (Colombia, Costa Rica, Mexico, Nicaragua)

T

tachigaliae (Forel, 1904) (Brazil, Peru)
   • Pseudomyrma bradleyi Enzmann, E.V., 1944
   • Pseudomyrma latinoda bradleyi Wheeler, W.M., 1942

tenuis (Fabricius, 1804) (Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Peru, Suriname)
   • Pseudomyrma ligniseca Smith, F., 1858
   • Pseudomyrma rufa Smith, F., 1877
   • Pseudomyrma tenuis andina Enzmann, E.V., 1944
   • Pseudomyrma tenuis paraensis Forel, 1912
   • Pseudomyrma tenuis pittieri Forel, 1906

tenuissimus (Emery, 1906) (Belize, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, El Salvador, French Guiana, Greater Antilles, Guatemala, Guyana, Honduras, Jamaica, Mexico, Nicaragua, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)
   • Pseudomyrma culmicola Forel, 1912

terminalis (Smith, F., 1877) (Brazil, Ecuador)
termitarius (Smith, F., 1855) (Bolivia, Brazil, Colombia, French Guiana, Grenada, Guyana, Lesser Antilles, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)
   • Pseudomyrma denticollis infusca Forel, 1908
   • Pseudomyrma elegans Smith, F., 1855
   • Pseudomyrma elegans breviceps Forel, 1912
   • Pseudomyrma nigriceps Smith, F., 1855

triplaridis (Forel, 1904) (Brazil, Colombia, Costa Rica, French Guiana, Guatemala, Guyana, Mexico, Panama, Suriname, Trinidad and Tobago, Venezuela)
   • Pseudomyrma triplaridis boxi Enzmann Enzmann, E.V., 1944
   • Pseudomyrma triplaridis boxi Wheeler, W.M. Wheeler, W.M., 1942

triplarinus (Weddell, 1850) (Argentina, Bolivia, Brazil, Colombia, Ecuador, Peru)
   • Pseudomyrma arborissanctae Emery, 1894
   • Pseudomyrma arborissanctae cordobensis Forel, 1914
   • Pseudomyrma arborissanctae ecuadoriana Enzmann, E.V., 1944
   • Pseudomyrma triplarina rurrenabaquensis Wheeler, W.M. & Mann, 1942

U

ultrix Ward, 1999 (Ecuador)
unicolor (Smith, F., 1855) (Bolivia, Brazil, Ecuador, French Guiana, Guyana, Mexico, Paraguay, Peru, Suriname)
   • Pseudomyrma atra Stitz, 1933
   • Pseudomyrma mutilloides Emery, 1890
   • Pseudomyrma unicolor anceps Santschi, 1925

urbanus (Smith, F., 1877) (Argentina, Bolivia, Brazil, Colombia, Costa Rica, Ecuador, French Guiana, Guyana, Panama, Paraguay, Peru, Suriname, Trinidad and Tobago, Venezuela)
   • Pseudomyrma chodati Forel, 1921
   • Pseudomyrma ogloblini Santschi, 1936

V

veneficus (Wheeler, W.M., 1942) (Mexico)
   • Pseudomyrma belti venifica Enzmann, E.V., 1944

venustus (Smith, F., 1858) (Bolivia, Brazil, Colombia, Panama, Peru)
   • Pseudomyrma agilis Smith, F., 1860
   • Pseudomyrma gracilis carapuna Mann, 1916
   • Pseudomyrma gracilis peruviana Enzmann, E.V., 1944
   • Pseudomyrma sedula Smith, F., 1877

veracruzensis Ward & Branstetter, 2022 (Mexico)
viduus (Smith, F., 1858) (Bolivia, Brazil, Colombia, Costa Rica, Cuba, Ecuador, French Guiana, Greater Antilles, Guyana, Honduras, Mexico, Suriname, Trinidad and Tobago)
   • Pseudomyrma caroli Forel, 1899
   • Pseudomyrma caroli sapii Forel, 1904
   • Pseudomyrma latinoda opacior Forel, 1904
   • Pseudomyrma triplaridis baileyi Wheeler, W.M., 1942
   • Pseudomyrma triplaridis biolleyi Enzmann, E.V., 1944
   • Pseudomyrma triplaridis tigrina Wheeler, W.M., 1942
   • Pseudomyrma triplaridis trigona Enzmann, E.V., 1944
   • Pseudomyrma ulei Forel, 1904

villosus Ward, 1989 (Brazil)
vinneni (Forel, 1906) (Brazil, Colombia, Peru, Venezuela)
vitabilis Ward, 1999 (Colombia)
voytowskii (Enzmann, E.V., 1944) (Peru)
voytowskii costaricensis (Enzmann, E.V., 1944) (Costa Rica)

W

weberi (Enzmann, E.V., 1944) (Guatemala)
wheeleri (Enzmann, E.V., 1944) (Peru)

Fossil Species

A

antiquus Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))
avitus Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

B

baros Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

C

coruscus Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

E

extinctus (Carpenter, 1930) (Florissant, Colorado, United States (Late Eocene))

M

macrops Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

N

nexilis Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

O

oryctus Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

P

prioris Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

S

saxulum LaPolla & Greenwlalt, 2015 (Kishenehn Formation shale, Montana, United States (Lutetian, Middle Eocene))
succinus Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

T

thecolor Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))

V

vicinus Ward, 1992 (Dominican amber, Dominican Republic (Burdigalian, Early Miocene))