Arnold (1915) observed these ants tending an unidentified Membracidae, one of just a few rare records of an army ant associating with honeydew producing insects.
Dorylus fulvus currently contains 11 subspecies and 7 junior synonyms. These taxa were described in the 1800's and early 1900's and the synonyms proposed in the early 1900's, with essentially no taxonomic consideration since. Of the 11 subspecies, 3 were described from workers, 4 were based on workers and males and 4 on males alone. This makes it almost impossible to taxonomically compare these taxa with each other, and makes the original descriptions essentially worthless. The true identity of these forms, and their relationships to each other, will remain uncertain until a full revision has been undertaken.
Photo Gallery
Dorylus fulvus worker, Israel. Photo by Eran Schnecke.
Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
Estimated Abundance
Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species.
homalinus. Cosmaecetes homalinus Spinola, 1851b: 55 (w.) Brazil (Pará).
Type-material: holotype worker.
Type-locality (in error): Brazil: Pará, 1846 (Ghiliani).
[Note: Spinola, 1853b: 71, says that he found three workers in M. Rambur’s collection that were exactly the same as the holotype, but from Algeria. Emery, 1893h: 1, confirms that homalinus is from North Africa.]
Type-depository: MRSN.
[Also described as new by Spinola, 1853: 71.]
[Misspelled as omalinus by Emery, 1893h: 1, Wheeler, W.M. 1911f: 160.]
Combination in Typhlopone: Smith, F. 1858b: 111;
combination in Dorylus: Dalla Torre, 1893: 11.
Status as species: Smith, F. 1858b: 111; Roger, 1861a: 45; Roger, 1863b: 20; Mayr, 1863: 457; Dalla Torre, 1893: 11.
spinolae. Labidus (Typhlopone) spinolae Shuckard, 1840c: 327 (w.) (no state data).
Type-material: holotype worker.
Type-locality: none given (G.H.K. Thwaites).
Type-depository: unknown (“in the collection of Mr Thwaites”.)
[Note: Smith, F. 1858b: 111, says that the single specimen was in “sugar casks from South America”. There is no such statement in the original description, but if the comment is correct then spinolae can not be a Dorylus species.]
Combination in Labidus: Westwood, 1842: 77;
combination in Typhlopone: Smith, F. 1858b: 111;
combination in Dorylus: Dalla Torre, 1893: 13.
Status as species: Smith, F. 1858b: 111; Roger, 1861a: 45; Roger, 1863b: 20; Mayr, 1863: 457; Dalla Torre, 1893: 13.
thwaitsii. Labidus (Typhlopone) thwaitsii Shuckard, 1840c: 326 (w.) (no state data).
Type-material: holotype(?) worker.
[Note: no indication of number of specimens is given.]
Type-locality: none given (G.H.K. Thwaites).
Type-depository: unknown (“in the collection of Mr Thwaites”.)
[Note: Smith, F. 1858b: 111, says that the specimen(s) were in, “sugar casks from South America”. There is no such statement in the original description, but if the comment is correct then thwaitsii can not be a Dorylus species.]
[Misspelled as twaitsi by Emery, 1910b: 13.]
Combination in Typhlopone: Smith, F. 1858b: 111;
combination in Dorylus: Dalla Torre, 1893: 13.
Status as species: Smith, F. 1858b: 111; Roger, 1861a: 45; Roger, 1863b: 20; Mayr, 1863: 457; Dalla Torre, 1893: 13.
Dalla Torre, K. W. von. 1893. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 7. Formicidae (Heterogyna). Leipzig: W. Engelmann, 289 pp. (page 10, Combination in Dorylus)
Emery, C. 1895l. Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zool. Jahrb. Abt. Syst. Geogr. Biol. Tiere 8: 685-778 (page 723, male described, Combination in D. (Typhlopone), Senior synonym of clausii, dahlbomii, shuckardi, spinolae and thwaitsii)
Westwood, J. O. 1839a. An introduction to the modern classification of insects; founded on the natural habits and corresponding organisation of the different families. Volume 2. Part XI. London: Longman, Orme, Brown, Green and Longmans, pp. 193-224. (page 219, fig. 86 worker described)
Barech G., M. Khaldi, S. Ziane, A. Zedam, S. Doumadji, M. Sharaf, and X. Espadaler. 2016. A first checklist and diversity of ants (Hymenoptera: Formicidae) of the saline dry lake Chott El Hodna in Algeria, a Ramsar Conservation Wetland. African Entomology 24(1): 143–152.
Begna D. 2007. Assesment of the effect of ant (Dorylus fulvus) on honeybee colony (A. mellifera) and their products in West & South-West Shewa zones, Ethiopia. Eth. J. Anim. Prod. 7(1): 12-26.
Borowiec L. 2014. Catalogue of ants of Europe, the Mediterranean Basin and adjacent regions (Hymenoptera: Formicidae). Genus (Wroclaw) 25(1-2): 1-340.
Cagniant, H. 1968. Liste preliminaire de fourmis forestieres d'Algerie. Resultats obtenus de 1963 a 1964. Bulletin de la Société d'Histoire Naturelle de Toulouse 104: 138-147
Cagniant, H. 2006. Liste actualisée des fourmis du Maroc. Myrmecologische Nachrichten 8: 193-200.
Cagniant, H.. "Liste actualisee des fourmis du Maroc (Hymenoptera: Formicidae)." Myrmecologische Nachrichten 8 (2006): 193-200.
Cagniant, H.. "Étude de quelques fourmis marocaines. Statistique provisoire des Formicidae du Maroc." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 53 (1964): 83-118.
Cagniant, H.. Les peuplements de fourmis des forêts algériennes: écologie, biocénotique, essai biologique. Universite de Toulouse, 1973.
Collingwood, C. A. and D. Agosti. 1996. Formicidae (Insects: Hymenoptera) of Saudi Arabia (Part 2) Fauna of Saudi Arabia 15: 300-385.
Crawley W. C. 1920. Ants from Mesopotamia and north-west Persia. Entomol. Rec. J. Var. 32: 162-166.
Dalla Torre, K. W.. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 7. Formicidae. Vol. 7. Leipzig: W. Engelmann, 1893.
Delye, G., and J. L. Bonaric. "Fourmis du sud Marocain." Etude de certains milieux du Maroc et de leux evolution recente RCP 249 (1973).
Derwesh A. I. 1965. A preliminary list of identified insects and some arachnids of Iraq. Ministry of Agriculture. Government Pressm Baghdad Iraq. Bulletin n 112: 128 pages.
Diame L., B. Taylor, R. Blatrix, J. F. Vayssieres, J. Y. Rey, I. Grechi, and K. Diarra. 2017. A preliminary checklist of the ant (Hymenoptera, Formicidae) fauna of Senegal. Journal of Insect Biodiversity 5(15): 1-16.
Donisthorpe H. 1918. A list of ants from Mesopotamia; with a description of a new species and a new variety. Entomologist's Record and Journal of Variation 30: 165-167.
Emery C. 1895. Die Gattung Dorylus Fab. und die systematische Eintheilung der Formiciden. Zoologische Jahrbücher. Abteilung für Systematik, Geographie und Biologie der Tiere 8: 685-778.
Emery, C.. "Exploration scientifique de la Tunisie. Zoologie. - Hyménoptères. Révision critique des fourmis de la Tunisie." Explor. Scient. De la Tunisie Zoll. Hym. (Folleto) Paris. Imp (1891): iii + 21 pp.
Field Museum Collection, Chicago, Illinois (C. Moreau)
Finzi B. 1939. Materiali zoologici dell'Eritrea raccolti da G. Müller durante la spedizione dell'Istituto Sieroterapico Milanese e conservati al Museo di Trieste. Parte III. Hymenoptera: Formicidae. Atti del Museo Civico di Storia Naturale di Trieste 14: 153-168.
Fisher B. L. 2004. Diversity patterns of ants (Hymenoptera: Formicidae) along an elevational gradient on Monts Doudou in southwestern Gabon. Memoirs of the California Academy of Sciences 28: 269-286.
Forel A. 1904. Miscellanea myrmécologiques. Rev. Suisse Zool. 12: 1-52.
Forel A. 1904. Note sur les fourmis du Musée Zoologique de l'Académie Impériale des Sciences à St. Pétersbourg. Ezheg. Zool. Muz. 8: 368-388.
Forel A. 1909. Études myrmécologiques en 1909. Fourmis de Barbarie et de Ceylan. Nidification des Polyrhachis. Bull. Soc. Vaudoise Sci. Nat. 45: 369-407.
Forel A. 1910. Glanures myrmécologiques. Ann. Soc. Entomol. Belg. 54: 6-32.
Forel A. 1910. Zoologische und anthropologische Ergebnisse einer Forschungsreise im westlichen und zentralen Südafrika ausgeführt in den Jahren 1903-1905 von Dr. Leonhard Schultze. Vierter Band. Systematik und Tiergeographie. D) Formicidae. Denkschriften der Medizinisch-Naturwissenschaftlichen Gesellschaft zu Jena 16: 1-30.
Forel, A.. "Fourmis de la faune méditerranéenne récoltées par MM. U. et J. Sahlberg." Revue Suisse de Zoologie 21 (1913): 427-438.
Goetsch, W. 1942. "Beiträge zur Biologie spanischer Ameisen." EOS (Revista española de entomología) 18: 175-241.
Hassanein A. H. M. 1990. Study of Species Dorylus fulvus Westwood (Subfamily: Dorylinae) in Libya. Journal of King Abdulaziz University , Educational Sciences 3, 1.
IZIKO South Africa Museum Collection
Kiran K., and C. Karaman. 2012. First annotated checklist of the ant fauna of Turkey (Hymenoptera: Formicidae). Zootaxa 3548: 1-38.
Kone M., S. Konate, K. Yeo, P. K. Kouassi, and K. E. Linsenmair. 2012. Changes in ant communities along an age gradient of cocoa cultivation in the Oumé region, central Côte dIvoire. Entomological Science 15: 324339.
Kugler J. 1988. The zoogeography of Israel. 9. The zoogeography of social insects of Israel and Sinai. Monographiae biologicae 62: 251-275.
Medler J. T. 1980: Insects of Nigeria - Check list and bibliography. Mem. Amer. Ent. Inst. 30: i-vii, 1-919.
Menozzi C. 1930. Formiche della Somalia italiana meridionale. Memorie della Società Entomologica Italiana. 9: 76-130.
Menozzi C. 1934. Reperti mirmecofaunistici raccolti dal Prof. L di Caporiacco nelle oasi di Cufra e in altre localita del deserto Libico. Atti della Societa dei Naturalisti e Matematici di Modena (6)13(65): 153-166
Ortiz, F. J., and A. Tinaut. "Introducción al conocimiento de las hormigas de la provincia de Almería." Publicaciones del Instituto de Estudios Almerienses. Boletín (Ciencias) 8 (1988): 223-231.
Rasheed M. T., I. Bodlah, A. G. Fareen, A. A. Wachkoo, X. Huang, and S. A. Akbar. 2019. A checklist of ants (Hymenoptera: Formicidae) in Pakistan. Sociobiology 66(3): 426-439.
Santschi, F.. "Quelques fourmis nord-africaines [FALTAN PÁGINAS]." Bulletin de la Société d' Histoire naturelle de l' Afrique du Nord 17 (1926): 229-236.
Tohme G. 1996. Formicidae. Etude de la diversité biologique n° 4 . Ministère de lAgriculture à Beyrouth (Eds.). P85-87.
Tohme G., and H. Tohme. 2014. Nouvelles liste des especes de fourmis du Liban (Hymenoptera, Formicoidea). Lebanese Science Journal 15(1): 133-141.
Weber N. A. 1952. Observations on Baghdad ants. Baghdad: The Trading and Printing Co., 30 pp.
Wheeler W. M. 1922. Ants of the American Museum Congo expedition. A contribution to the myrmecology of Africa. VIII. A synonymic list of the ants of the Ethiopian region. Bulletin of the American Museum of Natural History 45: 711-1004
Yeo K., S. Konate, S. Tiho, and S. K. Camara. 2011. Impacts of land use types on ant communities in a tropical forest margin (Oumé - Cote d'Ivoire). African Journal of Agricultural Research 6(2): 260-274.