Dorymyrmex pyramicus

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Dorymyrmex pyramicus
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Dolichoderinae
Tribe: Leptomyrmecini
Genus: Dorymyrmex
Species: D. pyramicus
Binomial name
Dorymyrmex pyramicus
(Roger, 1863)

Dorymyrmex pyramicus casent0173213 p 1 high.jpg

Dorymyrmex pyramicus casent0173213 d 1 high.jpg

Specimen Labels



Cuezzo and Guerrero (2011) - Worker Promesonotal profile continuous, strongly convex. Head, mesosoma, and legs reddish-yellow with gaster dark brown to black. Psammophore reaches the posterior margin of hypostoma. Queen Head subquadrate, maximum width at level of compound eyes; scape surpassing the posterior margin of head by no more than its maximum width; mandibles feebly striated with four teeth and two denticles on the masticatory margin; posterior margin of head straight. Forewing with only one large cubital cell. Male Head wider than long; posterior margin of head medially concave; scape long, surpassing the level of compound eyes, pygostyle poorly developed.

This species was described by Roger as Prenolepis pyramica from one worker collected in Bahia, Brazil, and transferred to Dorymyrmex by Mayr (1870). Unfortunately, Wheeler (1902) erroneously stated that Formica insana Buckley (1866) (Dorymyrmex insanus) was an “undoubtedly synonym” of D. pyramicus. Originally, Formica insana was described from Texas and southern states of the United States. Workers of D. insanus are concolorous black to dark brown, as Snelling (1973) says, differing from workers of D. pyramicus that are typically bicolored, as we describe above. Nevertheless, this mistake persisted, authors having considered D. pyramicus as a species with a very large distribution, from the south part of the United States throughout the Caribbean area to the north of Argentina. We only found one worker of D. pyramicus in Colombia, but there are bibliographic records that confirm its presence in this country (Fernandez and Sendoya 2004) . Apparently, D. insanus and D. pyramicus are only sympatric in Central America (Cuba and Guatemala) and in the north part of South America (Colombia, Venezuela, and Northern Brazil). Beside color, D. pyramicus and D. insanus can be differentiated by the shape of head in full-face view and the shape of the promesonotal profile (continuous in D. pyramicus, interrupted at its end in D. insanus).

Keys including this Species


Latitudinal Distribution Pattern

Latitudinal Range: 18.511028° to -38.85°.

Tropical South

Distribution based on Regional Taxon Lists

Neotropical Region: Argentina, Belize, Brazil (type locality), Colombia, Costa Rica, Cuba, Greater Antilles, Guatemala, Guyana, Lesser Antilles, Mexico, Suriname, Uruguay.

Distribution based on AntMaps


Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.


This species was found attending extrafloral nectaries of the plant Turnera subulata in northeastern Brazil (12°16′24″S 38°57′20″W / 12.27333°S 38.95556°W / -12.27333; -38.95556) Caatinga vegetation. Dorymyrmex pyramicus, classified as a dominant species, was a common attendant at the extrafloral nectaries.

Association with Other Organisms

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  • This species is a mutualist for the aphid Aphis lugentis (a trophobiont) (Jones, 1927; Saddiqui et al., 2019).
  • This species is a mutualist for the aphid Periphyllus negundinis (a trophobiont) (Jones, 1927; Saddiqui et al., 2019).
  • This species is a host for the phorid fly Pseudacteon dorymyrmecis (a parasitoid) (Quevillon, 2018) (encounter mode primary; direct transmission; transmission outside nest).



Images from AntWeb

Dorymyrmex pyramicus casent0909739 h 1 high.jpgDorymyrmex pyramicus casent0909739 d 1 high.jpgDorymyrmex pyramicus casent0909739 d 1 high.jpgDorymyrmex pyramicus casent0909739 l 1 high.jpg
Lectotype of Dorymyrmex pyramicus alticonisWorker. Specimen code casent0909739. Photographer Z. Lieberman, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Dorymyrmex pyramicus casent0909740 h 1 high.jpgDorymyrmex pyramicus casent0909740 p 1 high.jpgDorymyrmex pyramicus casent0909740 d 1 high.jpgDorymyrmex pyramicus casent0909740 l 1 high.jpg
Syntype of Dorymyrmex pyramicus mesonotalisWorker. Specimen code casent0909740. Photographer Z. Lieberman, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Dorymyrmex pyramicus casent0909741 h 1 high.jpgDorymyrmex pyramicus casent0909741 p 1 high.jpgDorymyrmex pyramicus casent0909741 d 1 high.jpgDorymyrmex pyramicus casent0909741 l 1 high.jpg
Syntype of Dorymyrmex pyramicus rubricepsWorker. Specimen code casent0909741. Photographer Z. Lieberman, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Dorymyrmex pyramicus casent0909743 h 1 high.jpgDorymyrmex pyramicus casent0909743 p 1 high.jpgDorymyrmex pyramicus casent0909743 d 1 high.jpgDorymyrmex pyramicus casent0909743 l 1 high.jpg
Holotype of Dorymyrmex antillanusWorker. Specimen code casent0909743. Photographer Z. Lieberman, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Dorymyrmex pyramicus casent0909746 h 1 high.jpgDorymyrmex pyramicus casent0909746 p 1 high.jpgDorymyrmex pyramicus casent0909746 d 1 high.jpgDorymyrmex pyramicus casent0909746 l 1 high.jpg
Lectotype of Dorymyrmex pyramicus garbeiWorker. Specimen code casent0909746. Photographer Z. Lieberman, uploaded by California Academy of Sciences. Owned by MHNG, Geneva, Switzerland.
Dorymyrmex pyramicus casent0911531 h 1 high.jpgDorymyrmex pyramicus casent0911531 p 1 high.jpgDorymyrmex pyramicus casent0911531 d 1 high.jpgDorymyrmex pyramicus casent0911531 l 1 high.jpg
Syntype of Dorymyrmex pyramicus guyanensisWorker. Specimen code casent0911531. Photographer Z. Lieberman, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Dorymyrmex pyramicus casent0911532 h 1 high.jpgDorymyrmex pyramicus casent0911532 p 1 high.jpgDorymyrmex pyramicus casent0911532 d 1 high.jpgDorymyrmex pyramicus casent0911532 l 1 high.jpg
Syntype of Dorymyrmex pyramicus nigriventrisWorker. Specimen code casent0911532. Photographer Z. Lieberman, uploaded by California Academy of Sciences. Owned by NHMB, Basel, Switzerland.
Dorymyrmex pyramicus castype00448-01 h 1 high.jpgDorymyrmex pyramicus castype00448-01 p 1 high.jpgDorymyrmex pyramicus castype00448-01 d 1 high.jpgDorymyrmex pyramicus castype00448-01 l 1 high.jpg
Syntype of Dorymyrmex pyramicus albemarlensisWorker. Specimen code castype00448-01. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Dorymyrmex pyramicus castype00448-02 h 1 high.jpgDorymyrmex pyramicus castype00448-02 p 1 high.jpgDorymyrmex pyramicus castype00448-02 d 1 high.jpgDorymyrmex pyramicus castype00448-02 l 1 high.jpg
Syntype of Dorymyrmex pyramicus albemarlensisWorker. Specimen code castype00448-02. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.


The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • pyramicus. Prenolepis pyramica Roger, 1863a: 160 (w.) BRAZIL. Emery, 1888c: 362 (m.); Gallardo, 1916a: 56 (q.); Wheeler, G.C. & Wheeler, J. 1951: 183 (l.); Goni, et al. 1983: 366 (k.). Combination in Dorymyrmex: Mayr, 1870b: 947; in D. (Conomyrma): Forel, 1913l: 244; in Conomyrma: Kusnezov, 1952g: 430; in Dorymyrmex: Shattuck, 1992c: 85. Current subspecies: nominal plus albemarlensis, alticonis, garbei, guyanensis, mesonotalis, nigriventris, peruvianus, rubriceps. See also: Emery, 1895c: 331; Snelling, R.R. 1973b: 4; Cuezzo & Guerrero, 2011: 19.

Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.



Cuezzo and Guerrero (2011) - (n = 10): HL: 0.78–0.88; HW: 0.73–0.78; EL 0.20–0.25; EW: 0.1–0.13; SL: 0.78–0.85; WL: 1.18–1.20; CI: 89–94; SI: 97–100; REL: 26–29; OI: 50–52; TLI: 137–152.

Head, mesosoma, and petiole concolorous reddish yellow; gaster always darker than the rest of the body, frequently dark brown to black. Whitish and sparse pubescence covering all body tagma. Head: posterior margin of head feebly emarginated medially. Psammophore with short hairs forming a triangle; the hairs in the top line are dispose near to the foramen magnum and do not reach the oral cavity. Upper seta line of psammophore close to anterior margin of foramen magnum. Mesosoma: pronotum with two subdecumbent short setae. Promesonotal profile strongly convex. Mesonotum not angulated, as in Dorymyrmex insanus. Propodeal tubercle well developed and directed dorsally. Declivitous face of propodeum, straight. Metasoma: petiolar scale pointing dorsally.


Cuezzo and Guerrero (2011) - (n = 2): HL: 1.15–1.18; HW: 1.18–1.2; EL: 0.33–0.4; EW: 0.13–0.15; IOD: 0.73; SL: 0.93–0.95; WL: 1.93–1.95; CI: 102-103; SI: 80-81.

Color and pubescence as in worker. Head: subquadrate; scape surpassing the posterior margin of head by no more than its maximum width; mandibles feebly striated, four teeth and two denticles on the masticatory margin; posterior margin of head straight; external margin of compound eye included in head surface in frontal view; ocelli hyaline, close to the posterior margin of head. Mesosoma: parapsidal furrows not well developed but parallels, axilla not divided medially. Anepisternum incompletely separated from katepisternum by a short suture. Forewing with only one close cubital cell; radial cell open. Metasoma: petiolar scale low, stout, and rounded apically.


Cuezzo and Guerrero (2011) - (n = 2): HL: 0.6–0.64; HW: 0.7–0.75; EL: 0.2–0.26; EW: 0.13–0.14; SL: 0.34; WL: 1.48–1.50.

Body color similar to worker and queen. Head: subquadrate, wider than long; lateral side of clypeus feebly projected forward; mandibles thin, with four teeth, apical tooth more than twice longer than the others; scape long, surpassing posterior margin of compound eyes. Mesosoma: parapsidal furrows present and parallel, axilla not divided medially; forewing with one close radial cell and no cubital nor discoidal cell. Hindwing with two basal cells. Metasoma: petiole stout and low, directed dorsally, ventral process round, feebly developed. Pygostyle poorly developed; gonystylus stout covered with few erect setae; digitus short and no cuspis. Aedeagus with serrate ventral border.


  • 2n = 18, karyotype = 14M+2SM+2A (Uruguay) (Goni et al., 1983) (as Conomyrma pyramica).


Cuezzo and Guerrero (2011) - The name of pyramicus refers to the typical tubercle or cone on propodeum present in all species of Dorymyrmex, giving to the propodeal angle an appearance of pyramid. Several Dorymyrmex species are known as “pyramid ants.”


References based on Global Ant Biodiversity Informatics

  • Alayo D. P. 1974. Introduccion al estudio de los Himenopteros de Cuba. Superfamilia Formicoidea. Academia de Ciencias de Cuba. Instituto de Zoologia. Serie Biologica no.53: 58 pp. La Habana.
  • Beck D. E., D. M. Allred, W. J. Despain. 1967. Predaceous-scavenger ants in Utah. Great Basin Naturalist 27: 67-78
  • Bezdeckova K., P. Bedecka, and I. Machar. 2015. A checklist of the ants (Hymenoptera: Formicidae) of Peru. Zootaxa 4020 (1): 101–133.
  • Borgmeier T. 1923. Catalogo systematico e synonymico das formigas do Brasil. 1 parte. Subfam. Dorylinae, Cerapachyinae, Ponerinae, Dolichoderinae. Archivos do Museu Nacional (Rio de Janeiro) 24: 33-103.
  • Canepuccia A. D., F. Hidalgo, J. L. Farina, F. Cuezzo, and O. O. Iribarne. 2016. Environmental harshness decreases ant β-diversity between salt marsh and neighboring upland environments. Wetlands DOI 10.1007/s13157-016-0777-0.
  • Cardoso, D.C., T.G. Sobrinho and J.H. Schoereder. 2010. Ant community composition and its relationship with phytophysiognomies in a Brazilian Restinga. Insectes Sociaux 57:293-301
  • Castano-Meneses, G., M. Vasquez-Bolanos, J. L. Navarrete-Heredia, G. A. Quiroz-Rocha, and I. Alcala-Martinez. 2015. Avances de Formicidae de Mexico. Universidad Nacional Autonoma de Mexico.
  • Cividanes F. J., J. C. Barbosa, I. C. F. Martins, F. Pattaro, M. A. Nunes, R. Souza Santos. 2009. Diversity and spatial distribution of ground arthropods in agroecosystems. Bragantia, Campinas, 68(4): 991-1002.
  • Clemes Cardoso D., and J. H. Schoereder. 2014. Biotic and abiotic factors shaping ant (Hymenoptera: Formicidae) assemblages in Brazilian coastal sand dunes: the case of restinga in Santa Catarina. Florida Entomologist 97(4): 1443-1450.
  • Clemes Cardoso D., and M. Passos Cristiano. 2010. Myrmecofauna of the Southern Catarinense Restinga sandy coastal plain: new records of species occurrence for the state of Santa Catarina and Brazil. Sociobiology 55(1b): 229-239.
  • Cole A. C., Jr. 1942. The ants of Utah. American Midland Naturalist 28: 358-388.
  • Cole, A.C. 1936. An annotated list of the ants of Idaho (Hymenoptera; Formicidae). Canadian Entomologist 68(2):34-39
  • Cuezzo F. and Guerrero, R. J. 2011. The ant genus Dorymyrmex Mayr (Hymenoptera: Formicidae: Dolichoderinae) in Colombia. Psyche 2012, Article ID 516058 (doi:10.1155/2012/516058): 24 pp
  • Dattilo W. et al. 2019. MEXICO ANTS: incidence and abundance along the Nearctic-Neotropical interface. Ecology
  • Drose W., L. R. Podgaiski, C. Fagundes Dias, M. de Souza Mendonca. 2019. Local and regional drivers of ant communities in forest-grassland ecotones in South Brazil: A taxonomic and phylogenetic approach. Plos ONE 14(4): e0215310.
  • Emery C. 1890. Studii sulle formiche della fauna neotropica. Bull. Soc. Entomol. Ital. 22: 38-8
  • Emery C. 1894. Estudios sobre las hormigas de Costa Rica. Anales del Museo Nacional de Costa Rica 1888-1889: 45-64.
  • Emery C. 1896. Formiche raccolte dal dott. E. Festa nei pressi del golfo di Darien. Bollettino dei Musei di Zoologia ed Anatomia Comparata della Reale Università di Torino 11(229): 1-4.
  • Emery C. 1906. Studi sulle formiche della fauna neotropica. XXVI. Bullettino della Società Entomologica Italiana 37: 107-194.
  • Escalante Gutiérrez J. A. 1993. Especies de hormigas conocidas del Perú (Hymenoptera: Formicidae). Revista Peruana de Entomología 34:1-13.
  • Fernández F., E. E. Palacio, W. P. Mackay, and E. S. MacKay. 1996. Introducción al estudio de las hormigas (Hymenoptera: Formicidae) de Colombia. Pp. 349-412 in: Andrade M. G., G. Amat García, and F. Fernández. (eds.) 1996. Insectos de Colombia. Estudios escogidos. Bogotá: Academia Colombiana de Ciencias Exactas, Físicas y Naturales, 541 pp
  • Fernández, F. and S. Sendoya. 2004. Lista de las hormigas neotropicales. Biota Colombiana Volume 5, Number 1.
  • Field Museum Collection, Chicago, Illinois (C. Moreau)
  • Fontanela J.L. 1999. Mirmecofauna del Jardin Botanico Nacional. Cocuyo 8: 8-10
  • Fontenla J. L. 2005. Species of ants (Formicidae) recorded during the rapid biological inventory of the Zapata Peninsula, 8-15 September 2002. In: Kirkconnell P., A., D. F. Stotz, y / and J. M. Shopland, eds. 2005. Cuba: Península de Zapata. Rapid Biological Inventories Report 07. The Field Museum, Chicago
  • Fontenla J. L., and J. Alfonso-Simonetti. 2018. Classification of Cuban ants (Hymenoptera: Formicidae) into functional groups. Poeyana Revista Cubana de Zoologia 506: 21-30.
  • Fontenla Rizo J. L. 1993. Composición y estructura de comunidades de hormigas en un sistema de formaciones vegetales costeras. Poeyana. Instituto de Ecología y Sistemática, Academia de Ciencias de Cuba 441: 1-19.
  • Fontenla Rizo J. L. 1993. Mirmecofauna de Isla de la Juventud y de algunos cayos del archipielago cubano. Poeyana. Instituto de Ecologia y Sistematica, Academia de Ciencias de Cuba 444:1-7.
  • Fontenla Rizo J. L., and L. M. Hernández. 1993. Relaciones de coexistencia en comunidades de hormigas en un agroecosistema de caña de azúcar. Poeyana. Instituto de Ecología y Sistemática, Academia de Ciencias de Cuba 438: 1-16.
  • Forel A. 1897. Quelques Formicides de l'Antille de Grenada récoltés par M. H. H. Smith. Transactions of the Entomological Society of London. 1897: 297-300.
  • Forel A. 1908. Catálogo systemático da collecção de formigas do Ceará. Boletim do Museu Rocha 1(1): 62-69.
  • Forel A. 1908. Fourmis de Costa-Rica récoltées par M. Paul Biolley. Bulletin de la Société Vaudoise des Sciences Naturelles 44: 35-72.
  • Forel A. 1912. Formicides néotropiques. Part V. 4me sous-famille Dolichoderinae Forel. Mémoires de la Société Entomologique de Belgique. 20: 33-58.
  • Franco W., N. Ladino, J. H. C. Delabie, A. Dejean, J. Orivel, M. Fichaux, S. Groc, M. Leponce, and R. M. Feitosa. 2019. First checklist of the ants (Hymenoptera: Formicidae) of French Guiana. Zootaxa 4674(5): 509-543.
  • Gallego-Ropero M.C., R.M. Feitosa & J.R. Pujol-Luz, 2013. Formigas (Hymenoptera, Formicidae) Associadas a Ninhos de Cornitermes cumulans Kollar (Isoptera, Termitidae) no Cerrado do Planalto Central do Brasil. EntomoBrasilis, 6(1): 97-101.
  • Gregg, R.T. 1963. The Ants of Colorado.
  • Hess C. G. 1958. The ants of Dallas County, Texas, and their nesting sites; with particular reference to soil texture as an ecological factor. Field and Laboratory 26: 3-72. 
  • Kempf W. W. 1978. A preliminary zoogeographical analysis of a regional ant fauna in Latin America. 114. Studia Entomologica 20: 43-62.
  • Kempf, W.W. 1972. Catalago abreviado das formigas da regiao Neotropical (Hym. Formicidae) Studia Entomologica 15(1-4).
  • Kirkconnell P., A., D. F. Stotz, y / and J. M. Shopland, eds. 2005. Cuba: Península de Zapata. Rapid Biological Inventories Report 07. The Field Museum, Chicago.
  • Kusnezov N. 1953. La fauna mirmecológica de Bolivia. Folia Universitaria. Cochabamba 6: 211-229.
  • Kusnezov N. 1957. Die Solenopsidinen-Gattungen von Südamerika (Hymenoptera, Formicidae). Zoologischer Anzeiger 158: 266-280.
  • Kusnezov N. 1963. Zoogeografia de las hormigas en sudamerica. Acta Zoologica Lilloana 19: 25-186
  • Kusnezov, N.. "Lista de las hormigas de Tucumán con descripción de dos nuevos géneros (Hymenoptera, Formicidae)." Acta Zoologica Lilloana 13 (1953): 327-339.
  • La Rivers I. 1968. A first listing of the ants of Nevada. Biological Society of Nevada, Occasional Papers 17: 1-12.
  • Lavigne R., and T. J. Tepedino. 1976. Checklist of the insects in Wyoming. I. Hymenoptera. Agric. Exp. Sta., Univ. Wyoming Res. J. 106: 24-26.
  • Llyod, Morrison W. 2002. Island bigeography and metapopulation dynamics of Bahamianan ants. Journal of Biogeography. 29:387-394.
  • Lopez, A. S., M. Vasquez-Bolanos, and G. A. Q. Rocha. 2015. Hormigas (Hymenoptera: Formicidae) del Cerro de la Culebra, Arandas, Jalisco, Mexico. Dugesiana 19: 151-155.
  • Lutinski J. A., B. C. Lopes, and A. B. Morais. 2013. Diversidade de formigas urbanas (Hymenoptera: Formicidae) de dez cidades do sul do Brasil. Biota Neotrop. 13(3): 332-342.
  • Mallis A. 1941. A list of the ants of California with notes on their habits and distribution. Bulletin of the Southern California Academy of Sciences 40: 61-100. 
  • Mann W. M. 1916. The Stanford Expedition to Brazil, 1911, John C. Branner, Director. The ants of Brazil. Bulletin of the Museum of Comparative Zoology 60: 399-490
  • Mann W. M. 1920. Additions to the ant fauna of the West Indies and Central America. Bulletin of the American Museum of Natural History 42: 403-439.
  • Menozzi C. 1935. Spedizione del Prof. Nello Beccari nella Guiana Britannica. Hymenoptera-Formicidae. Redia. 21: 189-203.
  • Morrison L. W. 1998. A review of Bahamian ant (Hymenoptera: Formicidae) biogeography. Journal of Biogeography 25: 561-571.
  • Morrison, Lloyd W. 1998. The Spatiotemporal Dynamics of Insular Ant Metapopulations. Ecology. 79(4):1135-1146.
  • Morrison, Lloyd. 2006. The Ants of Small Bahamian Cays. Bahamas Naturalist & Journal of Science. 1(2):27-32.
  • Nascimento Santos M., J. H. C. Delabie, and J. M. Queiroz. 2019. Biodiversity conservation in urban parks: a study of ground-dwelling ants (Hymenoptera: Formicidae) in Rio de Janeiro City. Urban Ecosystems
  • Osorio Rosado J. L, M. G. de Goncalves, W. Drose, E. J. Ely e Silva, R. F. Kruger, and A. Enimar Loeck. 2013. Effect of climatic variables and vine crops on the epigeic ant fauna (Hymenoptera: Formicidae) in the Campanha region, state of Rio Grande do Sul, Brazil. J Insect Conserv 17: 1113-1123.
  • Pergande T. 1893. On a collection of Formicidae from Lower California and Sonora, Mexico. Proc. Calif. Acad. Sci. (2) 4: 26-36.
  • Pignalberi C. T. 1961. Contribución al conocimiento de los formícidos de la provincia de Santa Fé. Pp. 165-173 in: Comisión Investigación Científica; Consejo Nacional de Investigaciones Científicas y Técnicas (Argentina) 1961. Actas y trabajos del primer Congreso Sudamericano de Zoología (La Plata, 12-24 octubre 1959). Tomo III. Buenos Aires: Librart, 276 pp.
  • Pires de Prado L., R. M. Feitosa, S. Pinzon Triana, J. A. Munoz Gutierrez, G. X. Rousseau, R. Alves Silva, G. M. Siqueira, C. L. Caldas dos Santos, F. Veras Silva, T. Sanches Ranzani da Silva, A. Casadei-Ferreira, R. Rosa da Silva, and J. Andrade-Silva. 2019. An overview of the ant fauna (Hymenoptera: Formicidae) of the state of Maranhao, Brazil. Pap. Avulsos Zool. 59: e20195938.
  • Radoszkowsky O. 1884. Fourmis de Cayenne Française. Trudy Russkago Entomologicheskago Obshchestva 18: 30-39.
  • Rivas-Arancibia, S. P., H. Carrillo-Ruiz, A. Bonilla-Arce, D. M. Figueroa-Castro, and A. R. Andres-Hernandez. 2014. Effect of disturbance on the ant community in a semiarid region of central Mexico. Applied Ecology and Environmental Research 12: 703-716.
  • Smith M. R. 1935. A list of the ants of Oklahoma (Hymen.: Formicidae). Entomological News 46: 235-241.
  • Smith M. R. 1936. A list of the ants of Texas. Journal of the New York Entomological Society 44: 155-170.
  • Smith, Marion R. 1954. American Museum Novitates. Ants of the Bimini Island Group, Bahamas, British West Indies (Hymenoptera, Formicidae). 1671:1-16
  • Smith, Marion R. 1954. Ants of the Bimini Island Group, Bahamas, British West Indies. American Museum of Natural History. 1671. 1-16.
  • Talbot M. 1976. A list of the ants (Hymenoptera: Formicidae) of the Edwin S. George Reserve, Livingston County, Michigan. Great Lakes Entomologist 8: 245-246.
  • Van Pelt A. F. 1966. Activity and density of old-field ants of the Savannah River Plant, South Carolina. Journal of the Elisha Mitchell Scientific Society 82: 35-43.
  • Vittar, F. 2008. Hormigas (Hymenoptera: Formicidae) de la Mesopotamia Argentina. INSUGEO Miscelania 17(2):447-466
  • Vittar, F., and F. Cuezzo. "Hormigas (Hymenoptera: Formicidae) de la provincia de Santa Fe, Argentina." Revista de la Sociedad Entomológica Argentina (versión On-line ISSN 1851-7471) 67, no. 1-2 (2008).
  • Vásquez-Bolaños M. 2011. Lista de especies de hormigas (Hymenoptera: Formicidae) para México. Dugesiana 18: 95-133
  • Warren, L.O. and E.P. Rouse. 1969. The Ants of Arkansas. Bulletin of the Agricultural Experiment Station 742:1-67
  • Weber N. A. 1948. Studies on the fauna of Curaçao, Aruba, Bonaire and the Venezuelan islands: No. 14. Ants from the Leeward Group and some other Caribbean localities. Natuurwetenschappelijke Studiekring voor Suriname en de Nederlandse Antillen 5: 78-86.
  • Wheeler W. M. 1901. The compound and mixed nests of American ants. Part II. The known cases of social symbiosis among American ants. American Naturalist. 35: 513-539.
  • Wheeler W. M. 1905. The ants of the Bahamas, with a list of the known West Indian species. Bulletin of the American Museum of Natural History 21: 79-135.
  • Wheeler W. M. 1907. A collection of ants from British Honduras. Bulletin of the American Museum of Natural History 23: 271-277.
  • Wheeler W. M. 1913. The ants of Cuba. Bulletin of the Museum of Comparative Zoology 54: 477-505.
  • Wheeler, William Morton. 1911. Additions to the Ant-Fauna of Jamaica. Bulletin American Museum of Natural History. 30:21-29.
  • Wheeler, William Morton. 1911. Ants Collected in Grenada, W.I. by Mr. C. T. Brues. Bulletin of the Museum of Comparitive Zoology at Harvard College. 54(5):166-172.
  • Wheeler, William Morton. 1934. Some Ants From The Bahama Islands. Psyche. 41(4):230-232.
  • Whitcomb W. H., H. A. Denmark, A. P. Bhatkar, and G. L. Greene. 1972. Preliminary studies on the ants of Florida soybean fields. Florida Entomologist 55: 129-142.
  • Wilson E. O. 1958. Sympatry of the ants Conomyrma bicolor (Wheeler) and C. pyramica (Roger). Psyche (Cambridge) 64: 76.
  • Young J., and D. E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publication. Oklahoma Agricultural Experimental Station 71: 1-42.
  • Young, J. and D.E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publications of Oklahoma State University MP-71
  • Zolessi L. C. de, Y. P. Abenante, and M. E. de Philippi. 1988. Lista sistematica de las especies de Formicidos del Uruguay. Comun. Zool. Mus. Hist. Nat. Montev. 11: 1-9.
  • Zolessi L. C. de; Y. P. de Abenante, and M. E. Philippi. 1989. Catálogo sistemático de las especies de Formícidos del Uruguay (Hymenoptera: Formicidae). Montevideo: ORCYT Unesco, 40 + ix pp.
  • de Zolessi, L.C., Y.P. de Abenante and M.E. Philippi. 1987. Lista sistemática de las especies de formícidos del Uruguay. Comunicaciones Zoologicas del Museo de Historia Natural de Montevideo 11(165):1-9
  • de Zolessi, L.C., Y.P. de Abenante and M.E. Phillipi. 1989. Catalago Systematico de las Especies de Formicidos del Uruguay (Hymenoptera: Formicidae). Oficina Regional de Ciencia y Technologia de la Unesco para America Latina y el Caribe- ORCYT. Montevideo, Uruguay