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Temporal range: Bartonian, Middle to Late Eocene Baltic amber, Baltic Sea region
Drymomyrmex fuscipennis
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Formicinae
Genus: Drymomyrmex
Wheeler, W.M., 1915
Type species
Drymomyrmex fuscipennis
2 fossil species
(Species Checklist, Species by Country)

The taxonomomic placement of this genus has been uncertain but Ward et al. (2016) places it within the Formicinae (but unplaced to tribe).



This taxon is known from Baltic amber (Bartonian, Middle to Late Eocene).



The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • DRYMOMYRMEX [incertae sedis in Formicidae]
    • Drymomyrmex Wheeler, W.M. 1915h: 135. Type-species: †Drymomyrmex fuscipennis, by original designation.
    • [This name is sometimes misspelled as †Dryomyrmex, for example in Donisthorpe, 1920: 93 and Wheeler, W.M. 1929a: 12.]
    • Drymomyrmex junior synonym of Camponotus: Dlussky, 1997: 59.
    • Drymomyrmex revived from synonymy: Dlussky & Rasnitsyn, 2009: 1036.


  • Bolton, B. 1994. Identification guide to the ant genera of the world. Cambridge, Mass.: Harvard University Press, 222 pp. (page 50, Drymomyrmex in Formicinae, Camponotini)
  • Bolton, B. 2003. Synopsis and Classification of Formicidae. Mem. Am. Entomol. Inst. 71: 370pp (page 112, Drymomyrmex as junior synonym of Camponotus [This name is sometimes misspelled as Dryomyrmex, for example in Donisthorpe, 1920:93 and Wheeler, W.M. 1929a:12])
  • Dlussky, G. M. 1997b. Genera of ants (Hymenoptera: Formicidae) from Baltic amber. Paleontol. Zh. 31: 616-627 (page 623, Drymomyrmex as junior synonym of Camponotus)
  • Dlussky, G. M.; Fedoseeva, E. B. 1988. Origin and early stages of evolution in ants. Pp. 70-144 in: Ponomarenko, A. G. (ed.) Cretaceous biocenotic crisis and insect evolution. Moskva: Nauka, 232 pp. (page 77, Drymomyrmex in Formicinae, Camponotini)
  • Donisthorpe, H. 1943g. A list of the type-species of the genera and subgenera of the Formicidae. [part]. Ann. Mag. Nat. Hist. 11(10): 617-688 (page 641, Drymomyrmex in Formicinae, Camponotini)
  • Ward, P.S., Blaimer, B.B., Fisher, B.L. 2016. A revised phylogenetic classification of the ant subfamily Formicinae (Hymenoptera: Formicidae), with resurrection of the genera Colobopsis and Dinomyrmex. Zootaxa 4072 (3): 343–357 (doi:10.11646/zootaxa.4072.3.4).
  • Wheeler, W. M. 1915i [1914]. The ants of the Baltic Amber. Schr. Phys.-Ökon. Ges. Königsb. 55: 1-142 (page 135, Drymomyrmex in Camponotinae, Camponotini)
  • Wheeler, W. M. 1929a. The identity of the ant genera Gesomyrmex Mayr and Dimorphomyrmex Ernest André. Psyche (Camb.) 36: 1-12 (page 12, Drymomyrmex in Formicinae, Brachymyrmecini)