Formica cunicularia

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Formica cunicularia
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Formicinae
Tribe: Formicini
Genus: Formica
Species: F. cunicularia
Binomial name
Formica cunicularia
Latreille, 1798

Formica cunicularia casent0173175 profile 1.jpg

Formica cunicularia casent0173175 dorsal 1.jpg

Specimen labels

Synonyms
Evolutionary Relationships

Proformica longiseta

Polyergus rufescens

Polyergus samurai

Iberoformica subrufa

Formica cunicularia

Formica gerardi

Formica frontalis

Formica sanguinea

Based on Gomez, Lorite, P. et al. 2018. Note that this study was restricted to species near Formica gerardi.

Species very common throughout Europe, even in disturbed habitats; also reported from Morocco (Rigato & Toni, 2011) and Russia (Zryanin & Zryanina, 2007). In Russia it is found in steppe habitats and dry pine forests. Nests are usually without mounds but mounds occur in some situations such as in meadows. Pashaei Rad et al. (2018) found this species in Iran on parkland ground in a moderate rainfall area. In Greece, Borowiec & Salata (2021) found this species to be common in northern mainland provinces, moderately common in southern mainland, on islands noted only from Aegean Islands and Crete. In northern provinces it was noted mostly from pastures, agricultural areas and rural sites in tourist resorts. In Achaia, it was observed in a rest area around a large pine tree and in high-growing oak forest.

Photo Gallery

  • Formica cunicularia queen. Photo by Michal Kukla.

Identification

Seifert and Schultz (2009) - A member of the Formica rufibarbis group. The separation of F. cunicularia and Formica clara represents the most difficult discrimination problem within the Formica rufibarbis group because there is a deficiency of strongly discriminating structural characters.

The weak point is that intraspecific colour polymorphism and loss of pigmentation by light or storage media could possibly affect the reliability of the pigmentation characters PIGM and CONT, but just these two characters have the largest loadings (canonical correlations) in the DA. These loadings are 0.788 in PIGM1.4 and 0.391 in CONT1.4 but only 0.336 in EYE1.4 and 0.170 in nPN1.4, the two best structural discriminators. Another problem are the isolated West Mediterranean populations of F. cunicularia from Corsica, Sardinia and the Sierra Nevada which were all allocated in the DA to the F. cunicularia cluster but possibly represent a third species. Integrative approaches including DNA analysis could bring more clarity into this issue.

Collingwood (1979) - Ashy grey black with at least genae and mesopleural articulations reddish; often most of alitrunk and head may be reddish. Gula and occiput bare. Erect hairs normally absent on pronotum but occasionally one or two short erect hairs may be present on promesonotum, never on upper margin of scale. Length: 4.0-6.5 mm.

Keys including this Species

Distribution

Seifert and Schultz (2009) - Temperate, Ponto-south-Siberian and Submediterranean species of the West Palaearctic, occurring from southernmost England and Iberia to West Siberia (85° E). In northwestern Europe, it goes north to southern Sweden (58° N) but has not reached southern Finland so far. Having a planar to colline distribution in the northern parts of its range, it climbs up to 1800 m in the Alps, up to 2400 m in the Caucasus and up to 2000 m in the South Siberian Tarbagatay Mountains.

North Africa to South Scandinavia, Portugal to Urals (Collingwood 1979).

Latitudinal Distribution Pattern

Latitudinal Range: 59.4242205° to 32.48611°.

   
North
Temperate
North
Subtropical
Tropical South
Subtropical
South
Temperate

Distribution based on Regional Taxon Lists

Oriental Region: India, Pakistan.
Palaearctic Region: Albania, Andorra, Armenia, Austria, Azerbaijan, Belarus, Belgium, Bulgaria, Channel Islands, China, Croatia, Czechia, Denmark, Estonia, Finland, France (type locality), Georgia, Germany, Greece, Hungary, Iberian Peninsula, Iran, Italy, Kyrgyzstan, Latvia, Lithuania, Luxembourg, Mongolia, Montenegro, Morocco, Netherlands, Norway, Poland, Portugal, North Macedonia, Republic of Moldova, Romania, Russian Federation, Slovakia, Slovenia, Spain, Sweden, Switzerland, Türkiye, Ukraine, United Kingdom of Great Britain and Northern Ireland.

Distribution based on AntMaps

AntMapLegend.png

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
pChart

Habitat

Borowiec and Salata (2022) - Thermophilous species, noted from both deciduous and coniferous forests, gorges with deciduous forests, mountain pastures, roadsides in coniferous forests, rest area in village with stone walls, old quarry close to coniferous forests, beaches with sparse deciduous trees and in salines. Often observed in grasslands, gardens and parks in towns and tourist resorts. Most records are from low altitude but on sunny slopes of mountains can reach a height of 1550 m.

Biology

Nests in soil or under stones.

Seifert and Schultz (2009) - Moderately thermophilic. Preferred habitats are meagre and semidry grasslands on differing geological outcrop, more rarely extremely xerothermic grasslands and also open ruderal, rural or disturbed habitats, including road or railway verges. In contrast to F. rufibarbis more frequently occurring on loamy soils with more developed herb layer and less often invading the urban zone. Colony foundation usually by single gynes but also pleometrotic. Nests moderately populous, usually containing 1000 - 1800 workers, sometimes weakly polygynous, but polydomous colonies unknown (as in all members of the group). Usually inhabits simple soil nests, construction of high mounds of mineral soil in meagre grassland with higher herb layer regularly observed. Usually timid and fugitive, but populous nests with large workers may be very aggressive during nest defence. Not territorial. Foraging at surface temperatures of up to 50°C, mainly on open surfaces and in the herb layer, but not avoiding bushes and trees. Zoophagous, trophobiotic and nectarivorous. Low position in dominance hierarchies of ant communities, usually inferior to even Lasius niger (Linnaeus, 1758), whom it carefully evades thanks to superior walking speed and well-developed visual sense, thus enabling coexistence at long-term food sources. May snatch large prey items from L. niger by swift surprise attack. Favoured host species for several socially parasitic ant species. Alates occur 7 July ± 12 d [16 June, 1 August], n = 17 (Seifert 2007).

Collingwood (1979) - This is a common species throughout Western Europe, nesting under stones or in small earth mounds, colonising railway embankments, sun exposed borders of woodland, dry open pasture and sea cliffs. Each nest is separate and normally has only one queen. Its habits are mainly predaceous and scavenging. Alatae occur in July and August

Foraging/Diet

Formica cunicularia collect honeydew.

Novgorodova (2015b) investigated ant-aphid interactions of a dozen honeydew collecting ant species in Western Siberia pine and aspen-birch-pine forests (54°7´N, 83°06´E, 200 m, Novosibirsk) and mixed-grass-cereal steppes with aspen-birch groves (53°44´N, 78°02´E, 110 m, near Karasuk) in the Novosibirsk Region and coniferous forests in the northeastern Altai (north end of Lake Teletskoe, 51°48´N, 87°17´E, 434 m). All of the ants studied had workers that showed high fidelity to attending particular aphid colonies, i.e, individual foragers that collect honeydew tend to return to the same location, and group of aphids, every time they leave the nest. F. cunicularia honeydew collecting activities also showed some other specialization but this was dependent on colony size. Smaller colonies (hundreds of workers) did not specialize. Larger colonies (>1,000 workers), during the summer months when the aphids and ants were most active, had individual foragers that specialized on either collecting honeydew, guarding, i.e., protecting aphids from competitors, transporting honeydew, or scouting for new aphid colonies. Some individuals did not specialize and behaved like foragers from smaller colonies, while others would specialize by returning to a specific aphid colony but would as readily guard aphids as they would collect honeydew. F. cunicularia tended Chaitophorus populeti (Panzer) and Aphis craccivora Koch.

Guiliani et al. (2019) observed this species foraging on extrafloral nectaries of the invasive Reynoutria x bohemica (Polygonaceae) in Tuscany. The habitats examined were river banks and disturbed habitats.

Association with Other Organisms

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Other Insects

  • This species is a host for the ant Formica sanguinea (a slave maker) (Ruano et al., 2018; Seifert, 2018; de la Mora et al., 2021).
  • This species is a host for the ant Polyergus rufescens (a slave maker) (Romani et al., 2006; D'Ettorre et al., 2000; Mori et al., 2000; Visiccio et al., 2000; Visiccio et al., 2001; Romani et al., 2006; Seifert, 2018; Trager, 2013; de la Mora et al., 2021).
  • This species is a host for the ant Formica exsecta (a temporary parasite) (de la Mora et al., 2021; Seifert, 2018).
  • This species is a host for the ant Formica pratensis (a temporary parasite) (de la Mora et al., 2021; Seifert, 2018).
  • This species is a host for the ant Formica rufa (a temporary parasite) (de la Mora et al., 2021; Seifert, 2018).
  • This species is a xenobiont for the ant Lasius flavus (a xenobiont) in United Kingdom (Kanizsai et al., 2013; Morley, 1945) (Foreshore. Under stone).
  • This species is associated with the aphids Acyrthosiphon pisum, Acyrthosiphon rubi, Aphis acetosae, Aphis brotericola, Aphis craccivora, Aphis fabae, Aphis nasturtii, Aphis pomi, Aphis solanella, Aphis spiraecola, Brachycaudus tragopogonis, Chaitophorus albus, Chaitophorus populeti, Chaitophorus populialbae, Cinara boerneri, Cinara palaestinesis, Cinara pini, Macrosiphoniella pulvera, Macrosiphum rosae and Sipha maydis (Saddiqui et al., 2019 and included references).
  • This species is a mutualist for the butterfly Zizeeria knysna (Obregon et al. 2015).
  • This species is a host for the braconid wasp Elasmosoma luxemburgense (a parasitoid) (Quevillon, 2018) (encounter mode primary; direct transmission; transmission outside nest).
  • This species is a host for the braconid wasp Neoneurus vesculus (a parasitoid) (Quevillon, 2018) (encounter mode primary; direct transmission; transmission outside nest).
  • This species is a host for the eucharitid wasp Eucharis adscendens (a parasitoid) (Quevillon, 2018) (multiple encounter modes; direct transmission; transmission outside nest).

Trematoda

  • This species is a host for the trematode Dicrocoelium dendriticum (a parasite) in Germany (Hohorst & Graefe, 1961).
  • This species is a host for the trematode Dicrocoelium lanceatum (a parasite) in Armenia (Arakelian et al., 1997).

Flight Period

X X
Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Source: antkeeping.info.

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Life History Traits

  • Mean colony size: 1,100 (Beckers et al., 1989)
  • Foraging behaviour: solitary forager (Beckers et al., 1989)

Castes

Worker

  • Liu, C. et al. 2020. Ants of the Hengduan Mountains, Figure 31, Formica cunicularia.

Queen

Images from AntWeb

Formica cunicularia casent0173176 head 1.jpgFormica cunicularia casent0173176 profile 1.jpgFormica cunicularia casent0173176 profile 2.jpgFormica cunicularia casent0173176 dorsal 1.jpgFormica cunicularia casent0173176 label 1.jpg
Queen (alate/dealate). Specimen code casent0173176. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • cunicularia. Formica cunicularia Latreille, 1798: 40 (w.q.m.) FRANCE. Combination in F. (Serviformica): Forel, 1915d: 64. Junior synonym of rufibarbis: Walckenaer, 1802: 161; Dalla Torre, 1893: 209; Ruzsky, 1905b: 385; Forel, 1915d: 64; Emery, 1916b: 255; Emery, 1925b: 250. Revived from synonymy and status as species: Yarrow, 1954a: 231. Senior synonym of rubescens: Yarrow, 1954a: 231; Dlussky, 1967a: 73; Bernard, 1967: 296; Seifert & Schultz, 2009: 261; of fuscorufibarbis: Dlussky, 1967a: 73; Dlussky & Pisarski, 1971: 166; of glebaria: Bernard, 1967: 296; Boven, 1977: 164; Agosti & Collingwood, 1987a: 59; of glauca (and its junior synonyms caucasica, katuniensis, montivaga, montaniformis, volgensis): Atanassov & Dlussky, 1992: 267; of fuscoides: Arakelian, 1994: 94; Seifert & Schultz, 2009: 261.
  • fuscorufibarbis. Formica fusca var. fuscorufibarbis Forel, 1874: 54 (w.q.) SWITZERLAND. Combination in F. (Serviformica): Forel, 1915d: 63. Raised to species: Forel, 1906c: 189. Subspecies of glebaria: Bondroit, 1918: 50; of rufibarbis: Dalla Torre, 1893: 210; Stitz, 1939: 357; Novak & Sadil, 1941: 107. Junior synonym of rufibarbis: Bernard, 1967: 297; of cunicularia: Dlussky, 1967a: 73; Dlussky & Pisarski, 1971: 166. See also comment in Seifert, 2002b: 266.
  • rubescens. Formica fusca var. rubescens Forel, 1904f: 423 (w.) SWITZERLAND. [Unresolved junior primary homonym of rubescens Leach, above.] Emery, 1909b: 196 (q.); Wheeler, W.M. 1913f: 498 (m.). Subspecies of glebaria: Bondroit, 1918: 50; Boven, 1947: 188. Junior synonym of cunicularia: Yarrow, 1954a: 231; Dlussky, 1967a: 73; Bernard, 1967: 296; Seifert & Schultz, 2009: 261.
  • fuscoides. Formica (Serviformica) cunicularia subsp. fuscoides Dlussky, 1967a: 74 (w.q.m.) ARMENIA. Junior synonym of cunicularia: Arakelian, 1994: 94; Seifert & Schultz, 2009: 261.

Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.

Description

Worker

Seifert and Schultz (2009) - Medium-sized Serviformica species (CS 1.365 mm); head slightly elongated (CL / CW1.4 1.131); Scape moderately long SL / CS1.4 1.073; distance of lateral ocelli moderate (OceD / CS1.4 0.164); eyes rather large (EYE / CS1.4 0.301), petiole rather wide (PEW / CS1.4 0.468). Clypeus with sharp median keel and fine longitudinal micro-carinulae. Frontal triangle finely transversely rippled and with 30 - 60 short pubescence hairs. Eyes with microsetae of 7 - 13 μm maximum length. Total mean of unilateral setae numbers on different body parts predicted for a specimen with CS = 1.4 mm: pronotum 1.1, mesonotum 0.8, petiole scale dorsal of spiracle 0.25, flexor profile of hind tibia 0.3. Posterior margin and underside of head and dorsolateral metapleuron as a rule without setae. Ventral coxae with long setae, dorsum of gaster with scattered, moderately long setae. Dorsal mesonotum in lateral aspect broadly rounded. Metanotal depression in larger individuals relatively deep. Propodeal dome rounded in lateral view, basal profile sometimes concave and in smaller specimens often straight. Dorsal crest of petiole in frontal view bluntly angled in smaller specimens to broadly convex in larger specimens, in some of the large individuals with straight or weekly excavate median portion. Petiole scale in lateral aspect rather thin, with convex anterior and more straight posterior profile. Gaster with transverse microripples of small average distance (RipD 4.6 μm) and covered by dense silvery pubescence (sqPDG 3.1). Pubescence on head, mesosoma and petiole dense. Typical colour pattern: Head with exception of round reddish-yellowish spots on anterior genae, dorsal promesonotum, coxae and all appendages dark brown, gaster blackish brown. Other body parts more or less reddish-yellowish. Nests with much lighter specimens having whole mesosoma, coxae and petiole uniformly reddish and such with very dark specimens having the reddish pigmentation reduced to a very small spot on frontal margin of ventrolateral mesonotum; exceptionally completely dark specimens occur.

Borowiec and Salata (2022) - Large with strongly marked size-variation, HL: 1.174-1.680 (mean 1.421); HW: 0.937-1.349 (mean 1.127); SL: 1.222-1.730 (mean 1.449); EL: 0.397-0.495 (mean 0.444); ML: 1.86-2.56; MW: 0.77-1.16. Color. Head bicolor, clypeus and genae, yellowish to red, rest of surface brown to black, the pale genae sharply limited from the dark posterior part of head also when the area above the genae is lighter brown colored than frontal and occipital parts of head; this sharp border between the light anterior and dark posterior parts of the head is also visible in the color of the underside of the head; mesosoma variably colored, from uniformly yellowish red or red to mostly brown infuscate, often only upper parts of pronotum and mesonotum with obscure spots of diffused borders, these dark spots can also cover the sides of the mesosoma or almost the entire mesosoma is brown and only the intersegmental sutures are lighter; in rare melanistic forms whole mesosoma and petiolar scale are uniformly brown then head always bicolored with gena distinctly paler than dark frons and occipital part of head brown, gaster from yellowish brown to dark brown with transparent white posterior margin of tergites, often anterior slope of first tergite paper colored than dorsal surface of the tergite, antennae yellowish, apical 3-6 antennomeres often gradually infuscate; legs variable colored from uniformly yellowish red to mostly brown, coxa often darker than femora and tibiae, or coxa and femora darker than tibiae and tarsi. Head. 1.2- 1.3 times longer than wide, in front of eyes softly converging anterad, behind eyes softly rounded, occipital margin straight to slightly convex. Clypeus with median keel, on the whole surface distinctly microsculptured, slightly trapezoidal, its anterior margin convex, sides convergent posterad, posterior margin truncate or shallowly concave in the middle, whole clypeal surface with moderately long and moderately dense appressed pubescence, a row of 10-12 moderately long setae close at the anterior margin and usually 8 long erected setae arranged in three rows 4-2-2, sometimes with additional very short, 2-3 erected setae, the longest anterior seta with length 0.190. Head distinctly microreticulate, appears dull and opaque, with short and sparse appressed pubescence not covering head surface, interocular are with two pairs and ocellar area with a pair of moderately long, erected yellow setae, sometimes ocellar area with 1-3 additional short erected setae, occasionally frons with only a pair or without setae, ventral side of head lacking erected setae. Scape moderately long, 1.2-1.4 times longer than width of head, thin, distinctly reaching beyond the occipital margin, distinctly, regularly widened from base to apex, its surface microreticulate, with short and dense appressed pubescence, erected setae absent. Funicular segments elongate, thin, first segment 1.5-1.6 times as long as second segment, the second segment approximately twice as long as wide, only slightly shorter than third segment, the rest of funicular segments clearly longer than broad. Eyes big, elongate oval, approximately 0.31 length of head. Mesosoma. Elongate in dorsal view distinctly constricted in the middle, 2.2-2.4 times as long as wide, dorsally and laterally distinctly microreticulated, surface indistinctly dull and opaque. In lateral view promesonotum convex, mesonotal groove moderately deep, propodeum strongly, regularly convex. Whole mesosomal surface covered with moderately long and moderately dense appressed pubescence not covering the mesosomal surface, pronotum sometimes without, usually with 4-10 (at most 15) short erected setae, the longest with length 0.087, mesonotum without or with 1-6 short erected setae, propodeum lacking erected setae. Waist and gaster. Petiolar scale broad, moderately thick in lateral view, apex rounded without setae. Gaster shorter than mesosoma, all tergites distinctly microreticulate, appears dull and opaque, covered with moderately long and dense appressed pubescence not completely covering surface of tergites. All tergites close to posterior margin with a row of setae, surface of tergites with short, sparse erected setae. Legs. Ventral surface of fore femora with row of 1-5 erected setae, sometimes without setae, of mid femora lacking erected setae or at most with single seta close to trochanter.

Type Material

Seifert and Schultz (2009) - Neotype worker labelled “FRA: 44.4947°N, 0.9597°E, Fumel, 120 m, in a garden, leg. Galkowski 2008.07.25” and “Neotype Formica cunicularia Latreille 1798, des. Seifert & Schultz 2009”; SMN Görlitz. In case of destruction or loss of the neotype specimen, a replacement neotype can be designated from a series of five mounted workers from the same nest series in SMN Görlitz and further five workers in Musee National d'Histoire Naturelle Paris.

Justification of the neotype fixation: A current search in the Latreille collection of MNHN Paris failed to detect a specimen interpretable as a primary type (J. Casevitz-Weulersse, pers. comm. 2008) and the literature gives no indication that a revisor ever has seen one. In order to establish an unambiguous standard for differentiation from similar species, we fixed a neotype in a sample from the terra typica which is in agreement with the traditional morphological conception of F. cunicularia.

Taxonomic Notes

Seifert and Schultz (2009) - West Mediterranean isolated populations: We do not at this time propose these deviating and isolated populations from Corsica, Sardinia and the Sierra Nevada as heterospecific from F. cunicularia. Differences to the continental population are a significantly narrower petiole (PEW / CS1.4 0.433) and slightly longer 1st tergite setae (GHL / CS1.4 7.24%). It seems to be the only species of the group from Corsica where Formica clara and Formica rufibarbis have not been reported so far.

Karyotype

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  • n = 27, 2n = 54 (France) (Hauschteck-Jungen & Jungen, 1976).

References

References based on Global Ant Biodiversity Informatics

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