Karyotype counts

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Karyotype Counts by Subfamily

  • Range of haploid karyotype numbers by subfamily.
Hapoid Karyotype Number.png


  • Range of diploid karyotype numbers by subfamily.
Dipoid Karyotype Number.png


  • Range of karyotype numbers by subfamily.
Subfamily Min. Haploid Max. Haploid Min. Diploid Max. Diploid
Amblyoponinae 24 24 48 48
Dolichoderinae 5 16 10 32
Dorylinae 15 23 30 46
Ectatomminae 10 23 20 46
Formicinae 6 27 16 54
Heteroponerinae 11 12 22 24
Myrmeciinae 1 41 2 81
Myrmicinae 4 30 8 60
Ponerinae 4 60 8 120
Proceratiinae 18 18 46 46

Species-level Karyotype Data

Explore Karyotype Data: All, Drilldown
Taxon Haploid Diploid Karyotype Locality Source Notes
Acanthomyrmex 22 Malaysia Goni et al., 1982
Acanthomyrmex 24 Sarawak Tjan et al., 1986
Acanthomyrmex 11 Malaysia Goni et al., 1982
Acanthomyrmex 11 22 Malaysia Imai et al., 1983
Acromyrmex ambiguus 38 14M + 12SM + 8ST + 4A Brazil de Castro et al., 2020
Acromyrmex ambiguus 38 2M+6SM+16ST+14A Uruguay Goñi et al., 1984
Acromyrmex ameliae 36 Brazil Cardoso et al. 2018; de Castro et al., 2020
Acromyrmex aspersus 38 8M+10SM+16ST+4A Brazil Teixeira et al., 2017
Acromyrmex balzani 38 12M+10SM+14ST+2A Brazil Barros et al., 2016
Acromyrmex balzani 38 12M+10SM+14ST+2A French Guiana Aguiar et al., 2020
Acromyrmex coronatus 19 38 12M+8SM+16ST+2A Brazil Barros et al., 2016
Acromyrmex crassispinus 38 Brazil Fadini & Pompolo, 1996
Acromyrmex crassispinus 38 12M + 20SM + 4ST + 2A Brazil de Castro et al., 2020
Acromyrmex disciger 38 10M+12SM+14ST+2A Brazil Barros et al., 2016
Acromyrmex echinatior 38 8M+6SM+14ST+10A Brazil Barros et al., 2016
Acromyrmex heyeri 38 2M+6SM+16ST+14A Brazil; Uruguay Goni et al., 1983; SantosColares et al., 1997
Acromyrmex hispidus 38 2M+6SM+16ST+14A Uruguay Goni et al., 1983
Acromyrmex lundii 38 10M + 14SM + 10ST + 4A Brazil de Castro et al., 2020
Acromyrmex niger 38 12M+14SM+10ST+2A Brazil Barros et al., 2016
Acromyrmex nigrosetosus 38 12M + 14SM + 10ST + 2A Brazil de Castro et al., 2020
Acromyrmex rugosus 38 16M+12SM+8ST+2A Brazil Barros et al., 2016
Acromyrmex striatus 22 20M + 2SM Brazil de Castro et al., 2020
Acromyrmex striatus 11 22 20M+2SM Brazil Cristiano et al., 2013
Acromyrmex subterraneus 38 Brazil Fadini & Pompolo, 1996
Acromyrmex subterraneus 38 14M + 18SM + 4ST + 2A Brazil de Castro et al., 2020
Acromyrmex subterraneus molestans 38 10M+10SM+16ST+2A Brazil Fadini & Pompolo, 1996; Teixeira et al., 2017
Acropyga 15 30 32 Indonesia; Malaysia; Sarawak Goni et al., 1982; Imai et al., 1985; Tjan et al., 1986
Acropyga acutiventris 28 29 Malaysia Imai et al., 1983 suggested Robertsonian polymorphism
Aenictus 15 30 Taiwan Hung et al., 1972 the first record for the subfamily
Aenictus brevicornis 24 India Imai et al., 1984
Aenictus laeviceps 22 Malaysia Imai et al., 1983
Allomerus decemarticulatus 28 18M+6SM+2A French Guiana Aguiar et al., 2020
Allomerus octoarticulatus 44 4SM+40A French Guiana Aguiar et al., 2020 as ''Allomerus octoarticulatus'' and ''Allomerus octoarticulatus demerarae''
Amblyopone 44 Australia Imai et al., 1977 as ''Amblyopone'' cf. ''fortis''
Amblyopone australis 24 48 Australia Imai et al., 1977
Anochetus 24 Malaysia Goni et al., 1982
Anochetus 30 India Imai et al., 1984
Anochetus 34 India Imai et al., 1984
Anochetus 17 34 Sarawak Tjan et al., 1986
Anochetus 19 Malaysia Goni et al., 1982
Anochetus altisquamis 30 12M+6SM+2ST+10T Brazil Santos et al., 2010
Anochetus graeffei 30 38 India; Indonesia Imai et al., 1984; Imai et al., 1985
Anochetus horridus 46 8M+4SM+34T French Guiana Mariano et al., 2011; Santos et al., 2010
Anochetus madaraszi 28 India Imai et al., 1984
Anochetus modicus 30 Indonesia Imai et al., 1985
Anochetus targionii 30 16M+2SM+2ST+10A French Guiana Aguiar et al., 2020
Anochetus yerburyi 30 India Imai et al., 1984
Anonychomyrma 16 Australia Crozier, 1977 as ''Iridomyrmex'' sp. ANIC-10
Anonychomyrma itinerans 8 16 Australia Crozier, 1968a as ''Iridomyrmex itinerans''
Anoplolepis gracilipes 17 34 India; Indonesia; Malaysia; Sarawak Goni et al., 1982; Tjan et al., 1986; Imai et al., 1983; Imai et al., 1984; Imai et al., 1985 as ''Anoplolepis longipes''
Aphaenogaster 30 Malaysia Goni et al., 1982
Aphaenogaster beccarii 30 46 India; Malaysia Imai et al., 1983; Imai et al., 1984
Aphaenogaster depilis 34 Tunisia Hauschteck-Jungen & Jungen, 1983
Aphaenogaster famelica 17 34 Japan Imai & Yosida, 1964; Imai, 1966; Imai, 1969; Imai, 1971
Aphaenogaster fulva 36 USA Crozier, 1977
Aphaenogaster gibbosa 11 16 17 34 4M+2SM+4ST+6T; 6M+4SM+8ST+16T Switzerland Hauschteck-Jungen & Jungen, 1983; Lorite et al., 2000; Palomeque et al., 1993b this karyotype is reported as fusion of two telocentric chromosomes
Aphaenogaster iberica 17 34 6M+4SM+8ST+16T Spain Palomeque et al., 1993a; Palomeque et al., 1993b; Lorite et al., 2000 ocurrence of supernumery chromosome segment
Aphaenogaster japonica 22 Japan Imai, 1969
Aphaenogaster lamellidens 38 USA Crozier, 1977; Taber & Cokendolpher, 1988
Aphaenogaster longiceps 45 46 Australia Imai et al., 1977 a metacentric pair is single and two telocentric corresponds the arms
Aphaenogaster miamiana 36 USA Crozier, 1977
Aphaenogaster osimenseis 16 32 Japan Imai & Yosida, 1964; Imai & Yosida, 1966; Imai, 1966; Imai, 1969; Imai, 1971
Aphaenogaster rudis 18 20 40 41 42 44 USA Crozier, 1977 suggested to be due supernumerary chromosome
Aphaenogaster sardoa 34 Tunisia Hauschteck-Jungen & Jungen, 1983
Aphaenogaster senilis 16 32 6M+6SM+20ST Spain Palomeque et al., 1993a; Palomeque et al., 1993b; Lorite et al., 2000
Aphaenogaster smythiesii 11 22 34 20SM+2M; 16M+18A India; Japan Imai, 1969; Imai, 1971; Imai et al., 1984
Aphaenogaster subterranea 11 22 Germany; Switzerland Hauschteck, 1962; Hauschteck-Jungen & Jungen, 1983
Aphaenogaster testaceopilosa 17 34 Croatia; Spain; Tunisia Hauschteck-Jungen & Jungen, 1983 Lorite and Palomeque 2010 states some issues about the identification/sampling location
Aphaenogaster tipuna 34 Taiwan Hung et al., 1972
Aphaenogaster treatae 42 USA Crozier, 1977
Apterostigma 20 24 32 6M+12SM+2A; 24M; 14M+6SM+10ST+2T Brazil; French Guiana; Panama Fadini & Pompolo, 1996; Mariano et al., 2011; Murakami et al., 1998
Apterostigma madidiense 23 14M+20SM+10ST+2A Brazil Barros et al., 2013
Apterostigma mayri 24 24M Panama Murakami et al., 1998
Apterostigma steigeri 22 20M+2SM Brazil Barros et al., 2013
Arnoldius pusillus 22 Australia Imai et al., 1977 as ''Bothriomyrmex pusillus''
Atta bisphaerica 22 12M+6SM+4A Brazil Fadini & Pompolo, 1996; Barros et al., 2014
Atta colombica 11 22 12M+6SM+4A Panama Murakami et al., 1998; Barros et al., 2014 ''Atta columbica'' is a writing mistake
Atta laevigata 22 12M+6SM+4A Brazil Fadini & Pompolo, 1996; Barros et al., 2014
Atta robusta 22 18M+2SM+2ST Brazil Barros et al., 2015; de Castro et al., 2020
Atta sexdens 22 12M+6SM+4A Brazil Fadini & Pompolo, 1996; SantosColares et al., 1997; Barros et al., 2014
Atta sexdens 22 18M + 4A Brazil de Castro et al., 2020
Atta sexdens 22 18M+2SM+2ST Brazil Aguiar et al., 2020
Azteca trigona 28 10M+4SM+14ST Brazil Cardoso et al., 2012b
Basiceros convexiceps 20 20M Brazil Mariano et al., 2013
Bothriomyrmex 11 Switzerland Hauschteck, 1963
Bothriomyrmex corsicus 11 Italy Hauschteck-Jungen & Jungen, 1983 as ''Bothriomyrmex gibbus''
Bothroponera 48 India Imai et al., 1984 near ''B. tesseronoda''
Bothroponera 52 India Imai et al., 1984 near ''B. tesseronoda''
Bothroponera rubiginosa 76 18M+58A India Imai et al., 1984
Brachymyrmex 18 Brazil Crozier, 1970b
Brachyponera 22 Malaysia Goni et al., 1982
Brachyponera 44 Indonesia Imai et al., 1985 near ''Pachycondyla obscurans'', now is ''Brachyponera obscurans''
Brachyponera chinensis 22 Japan Imai & Kubota, 1972 as ''Brachyponera sinensis''
Brachyponera lutea 16 Australia Imai et al., 1977
Brachyponera luteipes 11 22 2A+20M India; Japan Imai, 1969; Imai et al., 1984; Imai & Yosida, 1964
Calomyrmex 28 Australia Imai et al., 1977
Camponotus 18 Malaysia Imai et al., 1983
Camponotus 20 Sarawak Tjan et al., 1986
Camponotus 20 Taiwan Hung et al., 1972 'C. variegatus'' complex
Camponotus 20 20M India Imai et al., 1984 near ''C. infuscus''
Camponotus 32 Australia Imai et al., 1977
Camponotus 32 10M+22A India Imai et al., 1984 near ''C. variegatus''
Camponotus 34 India Imai et al., 1984 near ''C. variegatus''
Camponotus 34 10M+24A India Imai et al., 1984 near ''C. variegatus''
Camponotus 35 India Imai et al., 1984 near ''C. variegatus''
Camponotus 38 Australia Imai et al., 1977
Camponotus 38 Indonesia Imai et al., 1985
Camponotus 38 Malaysia Goni et al., 1982
Camponotus 38 Malaysia Imai et al., 1983
Camponotus 39 Indonesia Imai et al., 1985 mentined as a result of Robertsonian polymorphism
Camponotus 39 Malaysia Imai et al., 1983
Camponotus 40 Malaysia Imai et al., 1983
Camponotus 40 Sarawak Tjan et al., 1986
Camponotus 40 4M+36A+13B Brazil Mariano et al., 2001 variation observed for 2n=41-43 due B-chromsomes
Camponotus 44 Malaysia Goni et al., 1982
Camponotus 46 Australia Imai et al., 1977
Camponotus 48 Australia Imai et al., 1977
Camponotus 52 Malaysia Imai et al., 1983
Camponotus 10 20 Australia Imai et al., 1977
Camponotus 13 Malaysia Imai et al., 1983
Camponotus 14 Malaysia Imai et al., 1983
Camponotus 18 36 Malaysia Goni et al., 1982
Camponotus 19 Malaysia Goni et al., 1982
Camponotus 19 38 Australia Imai et al., 1977
Camponotus 20 Malaysia Imai et al., 1983
Camponotus 20 40 Malaysia Goni et al., 1982; Imai et al., 1983
Camponotus 20 40 Malaysia Goni et al., 1982
Camponotus 23 Australia Imai et al., 1977
Camponotus 23 46 Australia Imai et al., 1977
Camponotus 26 52 USA Crozier, 1970b 'C. impressus'' group
Camponotus 9 18 Japan Imai & Kubota, 1972 'Myrmamblys'' group
Camponotus 9 18 Japan Imai, 1966; Imai & Yosida, 1964
Camponotus aethiops 42 France; old Yugoslavia Hauschteck-Jungen & Jungen, 1983
Camponotus alii 42 Spain Hauschteck-Jungen & Jungen, 1983 Lorite and Palomeque 2010 states some issues about the identification/sampling location
Camponotus atriceps 20 40 4M+36A Brazil Mariano et al., 2001
Camponotus balzani 20 40 6M+34A Brazil Mariano et al., 2001
Camponotus barbatus taylori 24 24A India Imai et al., 1984 as ''Camponotus taylori''
Camponotus bonariensis 40 8M+32A Uruguay Goni et al., 1983
Camponotus cingulatus 40 4M+36A Brazil Mariano et al., 2001
Camponotus compressus 10 20 40 India; Tunisia Hauschteck-Jungen & Jungen, 1983; Kumbkarni, 1965 Lorite and Palomeque 2010 states some issues about the identification/sampling location
Camponotus consobrinus 23 46 Australia Imai et al., 1977
Camponotus crassisquamis 39 40 39A+1M; 40A India Imai et al., 1984 Lorite and Palomeque 2010 states the presence of heteromorphic chromosome in 2n=39
Camponotus crassus 10 20 20M Brazil Aguiar et al., 2016; Mariano et al., 2001
Camponotus cruentatus 18 20 12SM+5ST+3T Spain Hauschteck-Jungen & Jungen, 1983; Lorite et al., 2002b
Camponotus dolendus 20 20M India Imai et al., 1984
Camponotus femoratus 44 4ST+40T French Guiana Mariano et al., 2011
Camponotus festinus 19 38 Malaysia Imai et al., 1983
Camponotus foreli 17 34 Spain Hauschteck-Jungen & Jungen, 1983
Camponotus japonicus 13 14 26 Japan Imai, 1966; Imai, 1969; Imai & Kubota, 1972; Imai & Yosida, 1964 In Imai et al. 1966 as ''Camponotus'' sp.; Lorite and Palomeque 2010 states that this karyotype could be a mistake
Camponotus kiusiuensis 28 Japan Imai & Yosida, 1964; Imai, 1969; Imai, 1969
Camponotus lateralis 28 Switzerland Hauschteck, 1962
Camponotus ligniperda 14 28 Switzerland Hauschteck, 1961; Hauschteck-Jungen & Jungen, 1983
Camponotus mitis 20 20M India Imai et al., 1984
Camponotus mus 26 16SM+10A Uruguay Goni et al., 1983
Camponotus obscuripes 28 Japan Imai, 1969
Camponotus parius 40 40A India Imai et al., 1984
Camponotus pilicornis 50 Spain Hauschteck-Jungen & Jungen, 1983
Camponotus punctulatus 40 2SM+2ST+36A Uruguay Goni et al., 1983
Camponotus renggeri 40 Brazil Aguiar et al., 2016
Camponotus rufipes 20 39 40 2SM+2ST+36A; 3M+36A; 4M+36A Brazil; Uruguay Aguiar et al., 2016; Goni et al., 1983; Mariano et al., 2001 Lorite and Palomeque 2010 states the presence of heteromorphic chromosome in 2n=39
Camponotus rufoglaucus 18 Spain; Tunisia Hauschteck-Jungen & Jungen, 1983 Lorite and Palomeque 2010 states some issues about the identification/sampling location
Camponotus sericeiventris 40 4M+36A Brazil Mariano et al., 2001
Camponotus sericeus 44 2M+42A India Imai et al., 1984
Camponotus sylvaticus 20 40 Greece; Spain Hauschteck-Jungen & Jungen, 1983
Camponotus thraso 40 40A India Imai et al., 1984
Camponotus vagus 28 Switzerland Hauschteck, 1961
Camponotus variegatus 26 20M+6A India Imai et al., 1984
Camponotus vitiosus 9 18 Japan Imai, 1966; Imai, 1969; Imai & Kubota, 1972 In Imai 1966a as ''Camponotus'' sp. and in Imai 1969 as ''C. caryae'' group sp. but the authors correct the identification by ''C. tokioensis'' and the current valid name is ''C. vitiosus''
Cardiocondyla 40 Malaysia Goni et al., 1982
Cardiocondyla minutior 27 Japan Imai & Yamauchi, unpublished (Japanese Ant Image Database)
Cardiocondyla nuda 28 12M+16A India Imai & Kubota, 1981; Imai et al., 1984
Cardiocondyla wroughtonii 52 Imai & Yamauchi, unpublished, see Japanese Ant Image Database
Carebara 26 26M India Imai et al., 1984 as ''Oligomyrmex'' sp. 5
Carebara 34 Indonesia Imai et al., 1985 as ''Oligomyrmex'' sp.1
Carebara 36 Malaysia Goni et al., 1982 as ''Oligomyrmex'' sp. 1
Carebara 36 Sarawak Tjan et al., 1986 as ''Oligomyrmex'' sp.1
Carebara 38 Australia Imai et al., 1977 as ''Oligomyrmex'' sp. ANIC-6
Carebara 42 Indonesia Imai et al., 1985 as ''Oligomyrmex'' sp.2
Carebara 42 Sarawak Tjan et al., 1986 as ''Pheidologeton''
Carebara 44 Sarawak Tjan et al., 1986 as ''Oligomyrmex'' sp.2
Carebara 44 26M+18A India Imai et al., 1984 as ''Oligomyrmex'' sp. 2
Carebara 16 32 10M+6A India Imai et al., 1984 as ''Oligomyrmex'' sp. 4
Carebara asina 44 8M+36A India Imai et al., 1984 as ''Oligomyrmex asinus''
Carebara diversa 42 12M+30A India Imai et al., 1984 as ''Pheidologeton diversus''
Carebara sauteri 18 Taiwan Hung et al., 1972 as ''Oligomyrmex sauteri''
Cataglyphis bicolor 26 Israel; Tunesia Hauschteck-Jungen & Jungen, 1983
Cataglyphis iberica 26 Spain Hauschteck-Jungen & Jungen, 1983 as ''C. albicans''
Cataglyphis setipes 52 12M+40A India Imai et al., 1984 at figure subtitle it is mentioned 2n=54 but it may be a mistake since the formula is correct as ''desbribed in the results and the karyotype has ''26 chromosome pairs
Centromyrmex feae 44 14M+30A India Imai et al., 1984
Cephalotes pusillus 22 44 28M+16A Brazil Cristiano et al., 2017
Chelaner 22 Australia Imai et al., 1977
Chelaner 32 Australia Imai et al., 1977
Chelaner whitei 24 Australia Imai et al., 1977 as ''Chelaner whitei''
Colobostruma 22 22M Australia Imai et al., 1977
Colobostruma alinodis 11 22 22M Australia Crozier, 1968d
Crematogaster 24 Indonesia Imai et al., 1985
Crematogaster 24 Malaysia Goni et al., 1982; Imai et al., 1983
Crematogaster 26 Australia Imai et al., 1977 2n=39 was ''also observed in one worker
Crematogaster 26 Malaysia Goni et al., 1982; Imai et al., 1983
Crematogaster 26 Malaysia Imai et al., 1983
Crematogaster 26 Sarawak Tjan et al., 1986
Crematogaster 36 Malaysia Goni et al., 1982
Crematogaster 40 Japan Imai, 1969
Crematogaster 56 Indonesia Imai et al., 1985 2n=58 also observed due B-chromosome polymorphism
Crematogaster 12 24 Australia Imai et al., 1977
Crematogaster biroi 24 24M India Imai et al., 1984
Crematogaster brunnea 36 32M+4A India Imai et al., 1984
Crematogaster longispina 24 20M+4SM French Guiana Aguiar et al., 2020
Crematogaster nawai 26 Japan Imai & Yosida, 1964; Imai, 1966; Imai, 1969 as ''Crematogaster laboriosa''
Crematogaster rothneyi 50 4M+46A India Imai et al., 1984
Crematogaster subnuda 18 36 (31-36)M+(0-5)A India Imai et al., 1984 Karyotype formula as state by the authors
Cryptopone rotundiceps 12 Australia Imai et al., 1977
Cryptopone sauteri 14 28 Japan Imai & Yosida, 1964; Imai, 1969; Imai & Kubota, 1972
Cryptopone testacea 9 18 Malaysia; Sarawak Imai et al., 1983; Tjan et al., 1986
Cylindromyrmex brasiliensis 34 18M+16A Brazil Mariano et al., 2004b
Cyphomyrmex cornutus 22 10M+12SM French Guiana Mariano et al., 2011
Cyphomyrmex costatus 20 20M Panama Murakami et al., 1998
Cyphomyrmex rimosus 32 28M+4A Panama Murakami et al., 1998
Cyphomyrmex transversus 12 24 14M+6SM+4A French Guiana Aguiar et al., 2020
Dacetinops concinnus 16 Malaysia Imai et al., 1983
Diacamma 30 28M+2A India Imai et al., 1984
Diacamma 44 Malaysia Imai et al., 1983
Diacamma 58 Sarawak Tjan et al., 1986
Diacamma 66 Indonesia Imai et al., 1985
Diacamma 18 36 Malaysia Goni et al., 1982
Diacamma vagans 7 14 14M India Imai et al., 1984
Dinoponera australis 114 Brazil Santos et al., 2012
Dinoponera gigantea 82 Brazil Santos et al., 2012
Dinoponera lucida 106 18M+88A Brazil Mariano et al., 2004a Population polymorphism
Dinoponera lucida 57 114 Brazil Mariano et al., 2008 Population polymorphism
Dinoponera lucida 58 116 Brazil Mariano et al., 2008 Population polymorphism
Dinoponera lucida 59 118 Brazil Mariano et al., 2008 Population polymorphism
Dinoponera lucida 59 118 Brazil Santos et al., 2012
Dinoponera lucida 60 Brazil Santos et al., 2012
Dinoponera lucida 60 120 Brazil Mariano et al., 2008 Population polymorphism
Dinoponera quadriceps 92 Brazil Santos et al., 2012
Discothyrea 30 Indonesia Imai et al., 1985 near ''D. bryanti''
Doleromyrma 12 12M Australia Imai et al., 1977 'darwinianus'' group (as ''Iridomyrmex sp.ANIC-9)
Doleromyrma 7 14 14M Australia Imai et al., 1977 ''darwinianus'' group (as ''Iridomyrmex sp.ANIC-8)
Dolichoderus 18 Malaysia Goni et al., 1982
Dolichoderus attelaboides 38 2M+50SM+6A Brazil Santos et al., 2016
Dolichoderus bidens 18 6M+12SM Brazil Santos et al., 2016
Dolichoderus decollatus 38 6M+32SM Brazil Santos et al., 2016
Dolichoderus diversus 22 10M+12SM Brazil Santos et al., 2016
Dolichoderus imitator 38 6M+28SM+4A Brazil Santos et al., 2016
Dolichoderus imitator 46 6M+28SM+12A French Guiana Aguiar et al., 2020
Dolichoderus lutosus 10 4M+6SM Brazil Santos et al., 2016
Dolichoderus quadripunctatus 28 Japan Imai, 1969
Dolichoderus scabridus 14 28 Australia Crozier, 1966; Imai et al., 1977 in Crozier 1966 as ''Diceratoclinea scrabida''
Dolichoderus thoracicus 30 Indonesia; Malaysia; Sarawak Imai et al., 1983; Imai et al., 1985 as ''D. bituberculatus''
Dolichoderus thoracicus 33 Malaysia Imai et al., 1983 as ''D. bituberculatus''
Dolichoderus voraginosus 20 14M+6SM Brazil Santos et al., 2016
Dorymyrmex bicolor 13 26 USA Crozier, 1970b
Dorymyrmex flavus 26 USA Cokendolpher & Francke, 1984 as ''Conomyrma flava''
Dorymyrmex pulchellus 18 Brazil Crozier, 1968b; Crozier, 1970b as ''Dorymyrex'' sp. in Crozier 1968b
Dorymyrmex pyramicus 18 14M+2SM+2A Uruguay Goni et al., 1983 as ''Conomyrma pyramica''
Dorymyrmex thoracicus 9 Peru Crozier, 1970b
Echinopla 12 24 Malaysia Imai et al., 1983
Ectatomma brunneum 44 22M+22A Brazil Barros et al., 2008
Ectatomma edentatum 46 Brazil Barros et al., 2008
Ectatomma muticum 20 16M+4A Brazil Barros et al., 2008
Ectatomma permagnum 46 20M+26A Brazil Barros et al., 2008
Ectatomma tuberculatum 36 30M+6A Brazil Barros et al., 2008
Ectomomyrmex 38 16M+22A India Imai et al., 1984
Ectomomyrmex astutus 18 Indonesia Imai et al., 1985 as ''Pachycondyla astuta''
Ectomomyrmex astutus 22 Indonesia Imai et al., 1985 as ''Pachycondyla astuta''
Ectomomyrmex leeuwenhoeki 8 16 Malaysia Imai et al., 1983 as ''Pachycondyla leeuwenhoeki''
Epopostruma 10 20 20M Australia Crozier, 1968d
Erromyrma latinodis 70 India Imai et al., 1984 as ''Monomorium latinode''
Eurhopalothrix 18 Malaysia Imai et al., 1983 'E. procera'' group
Forelius mccooki 16 32 USA Crozier, 1970b; Crozier, 1975 as ''F. foetidus''
Formica 26 Crozier, 1975
Formica 27 Crozier, 1975 'Formica fusca'' group
Formica aquilonia 26 Finland Rosengren et al., 1980
Formica candida 6 52 Japan Imai, 1969 as ''F. picea''
Formica cinerea 27 54 Switzerland Hauschteck-Jungen & Jungen, 1976
Formica cunicularia 27 54 France Hauschteck-Jungen & Jungen, 1976
Formica dakotensis 26 Crozier, 1975
Formica exsecta 26 52 Switzerland Agosti & Hauschteck-Jungen, 1987; Hauschteck-Jungen & Jungen, 1976
Formica frontalis 52 Spain Lorite et al., 2002b; Lorite et al., 2004a
Formica fusca 27 54 Germany; Japan; Switerland Imai, 1969 ; Hauschteck-Jungen & Jungen, 1976
Formica gagates 27 54 France; old Yugoslavia Hauschteck-Jungen & Jungen, 1976
Formica gerardi 27 Spain Lorite et al., 1998a; Lorite et al., 2002b
Formica japonica 27 54 Japan Imai & Yosida, 1964; Imai, 1966; Imai, 1969; Crozier, 1975
Formica lemani 54 Switzerland Hauschteck-Jungen & Jungen, 1976
Formica lugubris 26 52 Finland; Switzerland Hauschteck-Jungen & Jungen, 1976; Rosengren et al., 1980
Formica montana 27 54 USA Hung, 1969
Formica obscuripes 26 USA Hung, 1969
Formica pergandei 26 Crozier, 1975
Formica picea 26 52 Switzerland Hauschteck-Jungen & Jungen, 1976
Formica polyctena 26 52 Finland; Switzerland Hauschteck-Jungen & Jungen, 1976; Rosengren et al., 1980
Formica pratensis 26 52 Finland; Switzerland Hauschteck-Jungen & Jungen, 1976; Rosengren et al., 1980
Formica pressilabris 26 Finland Rosengren et al., 1980
Formica reflexa 26 Crozier, 1975
Formica rufa 26 52 Switzerland Hauschteck-Jungen & Jungen, 1976
Formica rufibarbis 27 54 Finland; Switzerland Hauschteck-Jungen & Jungen, 1976; Rosengren et al., 1980
Formica sanguinea 26 52 Japan; Switzerland Imai & Yosida, 1964; Imai, 1966; Imai, 1969; Hauschteck-Jungen & Jungen, 1976
Formica subintegra 26 USA Hung, 1969
Formica transkaucasica 26 Finland Rosengren et al., 1980 as ''F. transkaucasica'' (incertae sedis)
Formica truncorum 26 52 Finland; Japan; Switzerland Imai & Yosida, 1964; Imai, 1969; Hauschteck-Jungen & Jungen, 1976; Rosengren et al., 1980
Formica truncorum 28 Finland Rosengren et al., 1980 Only one pupa analyzed was ''n=27
Formica ulkei 26 USA Hung, 1969
Formica uralensis 26 Finland Rosengren et al., 1980
Formica yessensis 26 52 Japan Imai, 1966; Imai, 1969
Formicoxenus chamberlini 28 USA Buschinger & Francoeur, 1983; Francoeur et al., 1985; Fischer, 1987 as ''Symmyrmica chamberlini''
Formicoxenus nitidulus 15 France Buschinger et al., 1980; Francoeur et al., 1985; Fischer, 1987
Formicoxenus provancheri 11 22 Canada Buschinger et al., 1980; Fischer, 1987 as ''Leptothorax provancheri''
Formicoxenus quebecensis 28 Canada Francoeur et al., 1985; Fischer, 1987
Gigantiops destructor 78 10M+66T French Guiana Mariano et al., 2011
Gnamptogenys 36 Malaysia Goni et al., 1982
Gnamptogenys 23 46 16M+30A Brazil Borges et al., 2004b
Gnamptogenys annulata 68 6M+62A Brazil Borges et al., 2004b
Gnamptogenys binghamii 22 Malaysia Imai et al., 1983
Gnamptogenys menadensis 42 Malaysia Goni et al., 1982; Imai et al., 1983
Gnamptogenys striatula 34 24M+10A Brazil Borges et al., 2004b
Harpagoxenus canadensis 18 Canada Buschinger et al., 1980; Fischer, 1987
Harpagoxenus sublaevis 20 40 Switzerland Buschinger et al., 1980; Hauschteck-Jungen & Jungen, 1983; Fischer, 1987
Heteroponera dolo 12 24 22M+2A Brazil Borges et al., 2004a
Heteroponera relicta 11 22 Australia Imai et al., 1977
Hypoponera 36 Malaysia Imai et al., 1983
Hypoponera 38 Australia Imai et al., 1977
Hypoponera 38 Malaysia Imai et al., 1983
Hypoponera 38 Malaysia Goni et al., 1982
Hypoponera 19 38 Australia Imai et al., 1977
Hypoponera confinis 38 Indonesia Imai et al., 1985
Hypoponera pruinosa 12 24 Indonesia; Malaysia; Sarawak Imai et al., 1983; Imai et al., 1985
Iberoformica subrufa 26 Spain Lorite et al., 2002b
Iridomyrmex 18 Australia Crozier, 1968a
Iridomyrmex 18 Australia Imai et al., 1977
Iridomyrmex 18 Malaysia Goni et al., 1982
Iridomyrmex 7 14 Australia Crozier, 1968a Lorite and Palomeque 2010 suggested that such karyotype is unusual in ''Iridomyrmex''
Iridomyrmex 9 18 Australia Crozier, 1968a
Iridomyrmex anceps 18 India Imai et al., 1984
Iridomyrmex anceps 48 Indonesia Imai et al., 1985 Lorite and Palomeque 2010 state that such karyotype is unusual in Dolichoderinae
Iridomyrmex bicknelli 9 18 Australia Crozier, 1968a as ''Iridomyrmex gracilis''
Iridomyrmex mattiroloi 9 18 Australia Crozier, 1968a
Iridomyrmex purpureus 9 18 Australia Crozier, 1968a; Crozier, 1968c; Imai et al., 1977 ''Iridomyrmex detectus'' in Crozier 1968a
Lasius alienus 28 Switzerland Hauschteck, 1962
Lasius alienus 15 30 Germany; Switzerland Hauschteck-Jungen & Jungen, 1983 n=14 was also reported, Lorite and Palomeque 2010 suggested that such karyotype is due Robertsonian polymorphism
Lasius brunneus 15 1M+3ST+11T Spain Lorite et al., 1998a; Lorite et al., 2002b Karyotype n=16 was reported in Lorite et al., 2002b and suggested that it is due B-chromosome
Lasius emarginatus 30 Switzerland Hauschteck, 1962
Lasius flavus 15 30 Switzerland Hauschteck, 1962; Hauschteck-Jungen & Jungen, 1983
Lasius fuliginosus 14 28 Italy; Switzerland Hauschteck, 1962; Hauschteck-Jungen & Jungen, 1983
Lasius nearcticus 15 30 USA Crozier, 1970b
Lasius niger 15 30 Japan; Spain; Switzerland Hauschteck, 1962; Hauschteck, 1963; Imai & Yosida, 1964; Imai, 1966; Imai, 1969; Imai & Kubota, 1972; Palomeque et al., 1990b, 1990c In Spain population in Palomeque et al. 1990 variation in the number of chromososmes were reported due B-chromosome
Lasius pallitarsis 28 USA Hung, 1969 as ''Lasius pallitarsus''
Lasius sakagamii 15 30 2M+28A Japan Yamauchi et al., 2001
Lasius talpa 30 Japan Imai & Yosida, 1964; Imai, 1966; Imai, 1969
Lasius umbratus 15 30 Switzerland; USA Hauschteck, 1962; Hung, 1969
Lepisiota 18 India Imai et al., 1984 near to ''Lepisiota sericea''
Lepisiota 18 India Imai et al., 1984 near to ''Lepisiota fergusoni''
Lepisiota capensis 18 India Imai et al., 1984 as ''Acantholepis lunaris''
Leptogenys 48 Malaysia Imai et al., 1983
Leptogenys 54 Sarawak Tjan et al., 1986
Leptogenys 15 30 India Imai et al., 1984 near ''Leptogenys peugueti''
Leptogenys borneensis 46 Malaysia Imai et al., 1983
Leptogenys diminuta 32 Indonesia Imai et al., 1985
Leptogenys diminuta 38 India; Malaysia Goni et al., 1982; Imai et al., 1983; Imai et al., 1984 in Gona et al. 1982 as ''Leptogenys'' sp.1
Leptogenys hysterica 26 India Imai et al., 1984
Leptogenys iridescens 46 Indonesia; Malaysia; Sarawak Imai et al., 1983; Imai et al., 1985
Leptogenys kraepelini 26 Indonesia Imai et al., 1985
Leptogenys myops 24 48 Indonesia; Malaysia; Sarawak Goni et al., 1982; Imai et al., 1983; Imai et al., 1985; Tjan et al., 1986 in Gona et al. 1982 as ''Leptogenys'' sp.2
Leptogenys peuqueti 52 India Imai et al., 1984 as ''Leptogenys minchini''
Leptogenys peuqueti 54 Indonesia Imai et al., 1985
Leptogenys processionalis 46 India Imai et al., 1984 as ''Leptogenys ocellifera''
Leptomyrmex erythrocephalus 12 Australia Imai et al., 1977
Leptothorax 12 Algeria Fischer, 1987
Leptothorax 12 Italy Fischer, 1987
Leptothorax 13 Croatia Fischer, 1987
Leptothorax 14 Canada Fischer, 1987
Leptothorax 15 Canada Loiselle et al., 1990
Leptothorax 16 Canada Loiselle et al., 1990
Leptothorax 16 Japan Imai, 1969
Leptothorax 17 USA Fischer, 1987
Leptothorax 17 USA Loiselle et al., 1990
Leptothorax 21 France Fischer, 1987
Leptothorax 22 France Fischer, 1987
Leptothorax acervorum 13 26 Canada; France; Germany; Sweden; Switzerland Fischer, 1987; Hauschteck-Jungen & Jungen, 1983; Loiselle et al., 1990
Leptothorax canadensis 16 Canada; USA Loiselle et al., 1990
Leptothorax canadensis 17 34 Canada; Germany; USA Buschinger et al., 1980; Fischer, 1987; Loiselle et al., 1990 possible variation due B chromosomes
Leptothorax canadensis 18 36 Canada; Germany; USA Fischer, 1987; Loiselle et al., 1990 possible variation due B chromosomes
Leptothorax canadensis 22 44 Canada; Germany; USA Fischer, 1987 possible variation due B chromosomes
Leptothorax canadensis 23 Canada; USA Loiselle et al., 1990 possible variation due B chromosomes
Leptothorax crassipilis 17 USA Francoeur, 1986; Loiselle et al., 1990
Leptothorax crassipilis 18 USA Francoeur, 1986; Loiselle et al., 1990
Leptothorax faberi 15 Canada Buschinger, 1982
Leptothorax faberi 16 Canada Fischer, 1987
Leptothorax goesswaldi 28 France; Switzerland Fischer, 1987 as ''Doronomyrmex goesswaldi''
Leptothorax gredleri 11 Germany; Switzerland Buschinger et al., 1980; Fischer, 1987; Loiselle et al., 1990
Leptothorax kutteri 23 24 25 France; Germany; Italy; Sweden; Switzerland Buschinger & Fischer, 1991; Douwes & Buschinger, 1983; Fischer, 1987 as ''Doronomyrmex kutteri''
Leptothorax muscorum 17 34 Canada; Germany; USA Buschinger et al., 1980; Fischer, 1987; Loiselle et al., 1990 possible variation due B chromosomes
Leptothorax muscorum 18 36 Canada; Germany; USA Fischer, 1987; Loiselle et al., 1990 possible variation due B chromosomes
Leptothorax muscorum 22 44 Canada; Germany; USA Fischer, 1987 possible variation due B chromosomes
Leptothorax pacis 26 France; Switzerland Buschinger & Fischer, 1991; Fischer, 1987 as ''Doronomyrmex pacis''
Leptothorax pocahontas 18 Canada Fischer, 1987; Buschinger & Heinze, 1993 as ''Doronomyrmex pocahontas''
Leptothorax retractus 17 Canada Francoeur, 1986; Loiselle et al., 1990
Leptothorax retractus 18 Canada Francoeur, 1986; Loiselle et al., 1990
Leptothorax sphagnicola 13 Canada Francoeur, 1986; Fischer, 1987; Loiselle et al., 1990 as ''Leptothorax sphagnicolus''
Linepithema fuscum 18 Peru Crozier, 1970b as ''Iridomyrmex pilifer'', reidentified as ''L. fuscum'' by Wild (2007)
Linepithema fuscum 18 Peru Crozier, 1970b as ''Iridomyrmex'' sp. nr. ''pilifer'', reidentified as ''L. fuscum'' by Wild (2007)
Linepithema humile 8 16 Australia; Spain Crozier, 1968a; Crozier, 1975; Lorite et al., 1996b; Lorite et al., 1998b as ''Iridomyrmex humilis''
Lioponera brevis 23 46 Australia Imai et al., 1977 as ''Cerapachys brevis''
Lophomyrmex 38 Sarawak Tjan et al., 1986
Lophomyrmex bedoti 38 India; Malaysia Imai et al., 1983; Imai et al., 1984
Lordomyrma 11 22 Malaysia Imai et al., 1983
Manica rubida 44 Switzerland Hauschteck, 1965; Hauschteck-Jungen & Jungen, 1983 as ''Myrmica rubida''
Mayaponera arhuaca 12 12M Brazil Mariano et al., 2007 as ''Pachycondyla arhuaca''
Mayaponera arhuaca 36 36A French Guiana Mariano et al., 2012 as ''Pachycondyla arhuaca''
Mayaponera constricta 30 30A Brazil Mariano et al., 2007 as ''Pachycondyla constricta''
Mayriella abstinens 18 Australia Imai et al., 1977
Megalomyrmex incisus 25 50 40M+10SM Brazil Cardoso et al., 2017
Meranoplus 22 Australia Imai et al., 1977
Meranoplus 22 Australia Crozier, 1970b 'Meranoplus hirsutus'' group
Meranoplus 10 Australia Crozier, 1966 'Meranoplus oceanicus'' group
Meranoplus bicolor 16 India; Indonesia Imai et al., 1984; Imai et al., 1985
Meranoplus minor 22 Australia Imai et al., 1977
Mesoponera 22 Malaysia Goni et al., 1982
Mesoponera 28 Malaysia Goni et al., 1982
Mesoponera 36 Malaysia Goni et al., 1982
Mesoponera rubra 20 Sarawak Tjan et al., 1986 as ''Pachycondyla rubra''
Mesoponera rubra 38 Malaysia Imai et al., 1983 as ''Pachycondyla rubra''
Mesoponera rubra 40 Malaysia Imai et al., 1983 as ''Pachycondyla rubra''
Mesoponera rubra 10 20 Indonesia Imai et al., 1985 as ''Pachycondyla rubra''
Messor 41 India Imai et al., 1984
Messor aciculatus 22 44 Japan Imai & Yosida, 1964; Imai, 1966; Imai, 1969
Messor barbarus 21 Spain Crozier, 1975; Lorite et al., 2002b
Monomorium 22 Australia Imai et al., 1977
Monomorium 22 Indonesia Imai et al., 1985
Monomorium 22 Malaysia Goni et al., 1982
Monomorium 34 India Imai et al., 1984
Monomorium 42 Australia Imai et al., 1977
Monomorium 11 Malaysia Imai et al., 1983
Monomorium 11 22 Malaysia Goni et al., 1982; Imai et al., 1983
Monomorium 11 22 Malaysia Imai et al., 1983
Monomorium indicum 21 India Imai et al., 1984
Monomorium indicum 22 India Imai et al., 1984
Monomorium minimum 22 USA Crozier, 1970b
Monomorium orientale 20 India Imai et al., 1984
Monomorium pharaonis 11 22 Japan; USA Smith & Peacock, 1957; Imai & Yosida, 1964
Monomorium rothsteini 22 Australia Imai et al., 1977 as ''Chelaner rothsteini''
Monomorium sahlbergi 16 India Imai et al., 1984
Monomorium subopacum 34 Spain Lorite et al., 2002b; Lorite et al., 2004b
Monomorium viridum 22 USA Crozier, 1970b as ''Monomorium viride''
Mycetomoellerius 9 18 16M+2SM Brazil Barros et al., 2014 as ''Trachymyrmex fuscus''
Mycetomoellerius holmgreni 10 20 20M Brazil Barros et al., 2018; Micolino et al., 2019; Micolino et al., 2020
Mycetomoellerius iheringi 10 20 18M + 2SM Brazil Micolino et al., 2020
Mycetomoellerius relictus 10 20 20M Brazil Barros et al., 2013; Micolino et al., 2020
Mycetomoellerius urichii 9 18 16M + 2SM Brazil Barros et al., 2013 as ''Mycetomoellerius fuscus''
Mycetophylax conformis 15 30 22M+8SM Brazil Cardoso et al., 2012a; Cardoso et al., 2014; Moura et al., 2020
Mycetophylax morschi 28 Brazil Moura et al., 2020
Mycetophylax morschi 13 26 18M+10SM+2A Brazil Cardoso et al., 2012a; Cardoso et al., 2014; Moura et al., 2020
Mycetophylax morschi 15 30 18M+6SM+2A Brazil Cardoso et al., 2012a; Cardoso et al., 2014; Moura et al., 2020
Mycetophylax simplex 18 36 20M+16SM Brazil Cardoso et al., 2012a; Cardoso et al., 2014
Mycocepurus 8 4M Panama Murakami et al., 1998
Mycocepurus goeldii 8 8M Brazil Barros et al., 2010
Myrmecia 68 Indonesia Imai et al., 1985
Myrmecia 6 12 8M+4ST Australia Imai et al., 1977 as cf. ''fulvipes''
Myrmecia arnoldi 53 Australia Hirai et al., 1996
Myrmecia arnoldi 55 Australia Hirai et al., 1996
Myrmecia arnoldi 57 Australia Hirai et al., 1996
Myrmecia arnoldi 59 Australia Hirai et al., 1996
Myrmecia arnoldi 60 Australia Hirai et al., 1996
Myrmecia arnoldi 60 Australia Meyne et al., 1995
Myrmecia arnoldi 64 Australia Hirai et al., 1996
Myrmecia arnoldi 66 Australia Hirai et al., 1996
Myrmecia banksi 10 Australia Imai et al., 1994; Hirai et al., 1994; Hirai et al., 1996; Taylor, 2015
Myrmecia banksi 9 Australia Imai et al., 1977; Imai et al., 1994; Taylor, 2015 putative mutant individual
Myrmecia brevinoda 84 Australia Imai et al., 1977
Myrmecia cephalotes 66 Australia Imai et al., 1977
Myrmecia chasei 47 Australia Meyne et al., 1995; Hirai et al., 1996
Myrmecia croslandi 1 2 Australia Taylor, 1991; Meyne et al., 1995; Hirai et al., 1994; Hirai et al., 1996; Imai et al., 1994 Complex ''pilosula''
Myrmecia croslandi 2 3 1M+1M+1A Australia Taylor, 1991; Meyne et al., 1995; Hirai et al., 1994; Hirai et al., 1996; Imai et al., 1992; Imai et al., 1994 Complex ''pilosula''
Myrmecia croslandi 2 3 1M+1SM +1M Australia Taylor, 1991; Meyne et al., 1995; Hirai et al., 1994; Hirai et al., 1996; Imai et al., 1992; Imai et al., 1994 Complex ''pilosula''
Myrmecia croslandi 2 4 2SM +1A+1A Australia Taylor, 1991; Meyne et al., 1995; Hirai et al., 1996; Imai et al., 1992; Imai et al., 1994 Complex ''pilosula''
Myrmecia forficata 50 44M+6A Australia Imai et al., 1977
Myrmecia forficata 51 Australia Imai et al., 1977
Myrmecia forficata 52 Australia Hirai et al., 1996
Myrmecia forficata 52 Australia Meyne et al., 1995
Myrmecia forficata 54 Australia Hirai et al., 1996
Myrmecia fulvipes 48 Australia Meyne et al., 1995; Hirai et al., 1996
Myrmecia fulvipes 50 Australia Hirai et al., 1996
Myrmecia fulvipes 30 60 Australia Imai et al., 1977
Myrmecia gulosa 38 Australia Imai et al., 1977; Meyne et al., 1995; Hirai et al., 1996
Myrmecia haskinsorum 12 Australia Imai et al., 1994 Complex ''pilosula''
Myrmecia haskinsorum 15 Australia Imai et al., 1994 Complex ''pilosula''
Myrmecia haskinsorum 17 Australia Imai et al., 1994 Complex ''pilosula''
Myrmecia haskinsorum 18 Australia Imai et al., 1994; Meyne et al., 1995; Hirai et al., 1996 Complex ''pilosula''
Myrmecia haskinsorum 20 Australia Imai et al., 1994 Complex ''pilosula''
Myrmecia haskinsorum 23 Australia Imai et al., 1994 Complex ''pilosula''
Myrmecia haskinsorum 24 Australia Imai et al., 1994 Complex ''pilosula''
Myrmecia imaii 6 Australia Imai et al., 1994 Complex ''pilosula''
Myrmecia imaii 8 8A Australia Imai et al., 1994; Hirai et al., 1994; Hirai et al., 1996 Complex ''pilosula''
Myrmecia mandibularis 27 56 Australia Hirai et al., 1996
Myrmecia mandibularis 28 56 Australia Hirai et al., 1996
Myrmecia michaelseni 27 Australia Hirai et al., 1996
Myrmecia occidentalis 64 Australia Hirai et al., 1996
Myrmecia pavida 44 Australia Hirai et al., 1996
Myrmecia piliventris 6 Australia Imai & Taylor, 1986; Imai et al., 1988a
Myrmecia piliventris 17 34 Australia Imai & Taylor, 1986; Imai et al., 1988a
Myrmecia piliventris 2 4 4ST Australia Imai & Taylor, 1986; Imai et al., 1988a
Myrmecia piliventris 32 64 Australia Imai & Taylor, 1986; Imai et al., 1988a
Myrmecia pilosula 10 Australia Imai et al., 1977; Crossland & Crozier, 1986; Crosland et al., 1988 as ''Myrmecia ruginoda'' or ''Ponera ruginoda''
Myrmecia pilosula 16 Australia Crossland & Crozier, 1986
Myrmecia pilosula 18 Australia Crosland et al., 1988
Myrmecia pilosula 19 Australia Hirai et al., 1996
Myrmecia pilosula 2 Australia Crosland et al., 1988; Imai et al., 1988b; Imai & Taylor, 1989
Myrmecia pilosula 20 Australia Crosland et al., 1988
Myrmecia pilosula 21 Australia Crosland et al., 1988
Myrmecia pilosula 22 Australia Crosland et al., 1988
Myrmecia pilosula 22 Australia Hirai et al., 1996
Myrmecia pilosula 23 Australia Crosland et al., 1988; Imai et al., 1988a
Myrmecia pilosula 23 Australia Hirai et al., 1996
Myrmecia pilosula 23 Australia Imai et al., 1994
Myrmecia pilosula 23 Australia Meyne et al., 1995
Myrmecia pilosula 24 Australia Crossland & Crozier, 1986
Myrmecia pilosula 24 Australia Crosland et al., 1988; Imai et al., 1988a
Myrmecia pilosula 24 Australia Imai et al., 1994
Myrmecia pilosula 25 Australia Crosland et al., 1988; Imai et al., 1988a
Myrmecia pilosula 26 Australia Crosland et al., 1988; Imai et al., 1988a
Myrmecia pilosula 26 Australia Imai et al., 1994
Myrmecia pilosula 27 Australia Crosland et al., 1988
Myrmecia pilosula 27 Australia Hirai et al., 1994
Myrmecia pilosula 27 Australia Imai et al., 1994
Myrmecia pilosula 30 Australia Crossland & Crozier, 1986; Crosland et al., 1988
Myrmecia pilosula 31 Australia Imai et al., 1977; Crossland & Crozier, 1986; Crosland et al., 1988; Imai et al., 1988a as ''Myrmecia ruginoda'' or ''Ponera ruginoda''
Myrmecia pilosula 32 Australia Imai et al., 1977; Crossland & Crozier, 1986; Crosland et al., 1988; Imai et al., 1988a as ''Myrmecia ruginoda'' or ''Ponera ruginoda''
Myrmecia pilosula 32 Australia Hirai et al., 1996
Myrmecia pilosula 32 Australia Meyne et al., 1995
Myrmecia pilosula 4 Australia Imai et al., 1988b; Imai & Taylor, 1989
Myrmecia pilosula 9 Australia Imai et al., 1977; Crossland & Crozier, 1986; Crosland et al., 1988 as ''Myrmecia ruginoda'' or ''Ponera ruginoda''
Myrmecia pilosula 1 Australia Crosland et al., 1988 Population from Tridbinbilla
Myrmecia pilosula 1 19 Australia Crosland et al., 1988
Myrmecia pilosula 10 Australia Crosland et al., 1988 Population from Lawson
Myrmecia pilosula 10 Australia Crosland et al., 1988 Population from Mt Victoria
Myrmecia pilosula 10 Australia Crosland et al., 1988 Population from Wentworth Falls
Myrmecia pilosula 11 Australia Crosland et al., 1988 Population from Lawson
Myrmecia pilosula 11 Australia Crosland et al., 1988 Population from Mt Victoria
Myrmecia pilosula 13 Australia Crosland et al., 1988 Population from Mt Victoria
Myrmecia pilosula 15 Australia Crosland et al., 1988 Population from Mt Victoria
Myrmecia pilosula 15 Australia Crozier, 1966
Myrmecia pilosula 16 Australia Crosland et al., 1988 Population from Piccadilly Circus
Myrmecia pilosula 5 Australia Crosland et al., 1988 Population from HMAS Albatross
Myrmecia pilosula 9 Australia Crosland et al., 1988 Population from Wentworth Falls
Myrmecia pyriformis 41 81 Australia Imai et al., 1977
Myrmecia simillima 35 70 Australia Hirai et al., 1996
Myrmecia tepperi 70 Australia Browning, 1987
Myrmecia vindex 74 Australia Hirai et al., 1996
Myrmecia vindex 76 Australia Hirai et al., 1996
Myrmecina 66 Indonesia Imai et al., 1985
Myrmecina americana 14 Crozier, 1975
Myrmecina graminicola 14 Hauschteck, 1965; Crozier, 1975
Myrmica lobicornis 24 Hauschteck-Jungen & Jungen, 1983
Myrmica rubra 48 Switzerland Hauschteck-Jungen & Jungen, 1983 as ''Myrmica laevinodis''
Myrmica rubra 23 46 Japan Imai, 1969
Myrmica ruginodis 48 Switzerland Hauschteck, 1965
Myrmica ruginodis 24 Switzerland Hauschteck-Jungen & Jungen, 1983
Myrmica sabuleti 46 Switzerland Hauschteck, 1965
Myrmica sabuleti 23 Switzerland Hauschteck-Jungen & Jungen, 1983
Myrmica scabrinodis 22 44 Switzerland Hauschteck, 1965
Myrmica schencki 23 46 Switzerland Hauschteck-Jungen & Jungen, 1983
Myrmica sulcinodis 24 Switzerland Crozier, 1975 2n=24 is probably a mistake
Myrmica sulcinodis 28 56 Switzerland Hauschteck, 1965; Hauschteck-Jungen & Jungen, 1983
Myrmicaria 44 Indonesia Imai et al., 1985
Myrmicaria 44 Malaysia Goni et al., 1982
Myrmicaria 44 Malaysia Goni et al., 1982; Imai et al., 1983
Myrmicaria 23 46 Malaysia Imai et al., 1983
Myrmicaria brunnea 44 24M+20A India Imai et al., 1984 The photo is the ''Myrmicaria brunnea subcarinata''
Myrmicocrypta 30 22M+2SM+6A French Guiana Aguiar et al., 2020
Myrmicocrypta 30 6M+10SM+14A French Guiana Mariano et al., 2011
Mystrium camillae 32 Sarawak Tjan et al., 1986
Neivamyrmex texanus 36 36M USA Taber & Cokendolpher, 1988
Neoponera 54 6M+48A Brazil Mariano et al., 2007 as ''Pachycondyla venusta'' group
Neoponera apicalis 36 28M+8A Brazil Mariano et al., 2007; Delabie et al., 2008 as ''Pachycondyla apicalis''
Neoponera apicalis 40 30M+10A Brazil Mariano et al., 2007; Delabie et al., 2008 as ''Pachycondyla apicalis''
Neoponera apicalis 68 44M+20A Brazil Mariano et al., 2007; Delabie et al., 2008 as ''Pachycondyla apicalis''
Neoponera carinulata 24 4M+20A Brazil Mariano et al., 2007 as ''Pachycondyla carinulata''
Neoponera crenata 26 2M+24A Brazil Mariano et al., 2006a; Mariano et al., 2007 as ''Pachycondyla crenata''
Neoponera curvinodis 26 4M+22A Brazil Mariano et al., 2007 as ''Pachycondyla subversa''
Neoponera curvinodis 28 22M+6A Brazil Mariano et al., 2007 as ''Pachycondyla subversa''
Neoponera goeldii 24 24A French Guiana Mariano et al., 2007 as ''Pachycondyla goeldii''
Neoponera inversa 30 20M+10A Brazil Mariano et al., 1999; Mariano et al., 2007 as ''Pachycondyla inversa''
Neoponera marginata 46 28M+18A Brazil Mariano et al., 2007 as ''Pachycondyla marginata''
Neoponera metanotalis 70 16M+54A Brazil Mariano et al., 2012 as ''Pachycondyla metanotalis''
Neoponera moesta 13 26 26A Brazil Mariano et al., 2006a; Mariano et al., 2007 as ''Pachycondyla mesonotalis''
Neoponera unidentata 12 12M Brazil Mariano et al., 2007 as ''Pachycondyla unidentata''
Neoponera venusta 48 26M+22A Brazil Mariano et al., 2007 as ''Pachycondyla venusta''
Neoponera verenae 42 30M+12A Brazil Mariano et al., 2007; Delabie et al., 2008
Neoponera verenae 58 Brazil Mariano et al., 2007; Delabie et al., 2008
Neoponera verenae 58 14M+44A Brazil Mariano et al., 2007; Delabie et al., 2008 Different karyotype reported in Mariano 2007
Neoponera verenae 60 Brazil Mariano et al., 2007; Delabie et al., 2008 Different karyotype reported in Mariano 2007
Neoponera verenae 62 14M+48A Brazil Mariano et al., 2007; Delabie et al., 2008
Neoponera verenae 64 12M+52A Brazil Mariano et al., 2007; Delabie et al., 2008
Neoponera villosa 34 12M+22A Brazil Mariano et al., 1999; Mariano et al., 2007 as ''Pachycondyla villosa''
Nothomyrmecia macrops 94 Australia Imai et al., 1990
Notoncus ectatommoides 22 44 Australia Imai et al., 1977
Nylanderia 16 Malaysia Goni et al., 1982
Nylanderia 16 Malaysia Imai et al., 1983
Nylanderia 26 Malaysia Goni et al., 1982
Nylanderia 28 Malaysia Goni et al., 1982
Nylanderia 30 Australia Imai et al., 1977
Nylanderia 30 Indonesia Imai et al., 1985
Nylanderia 30 Malaysia Imai et al., 1983
Nylanderia 15 30 30A India Imai et al., 1984 near ''P. yerbuyi''
Nylanderia 8 16 Malaysia Goni et al., 1982; Imai et al., 1983
Nylanderia indica 30 India Imai et al., 1984 as ''Paratrechina indica''
Nylanderia parvula 15 Crozier, 1975 as ''Paratrechina parvula''
Ochetellus glaber 14 28 Australia; Japan Imai, 1969; Imai & Yosida, 1964 as ''Iridomyrmex glaber''n Crozier 1968a and as ''Iridomyrmex itoi''n Imai and Yosida 1964
Odontomachus 44 Australia Imai et al., 1977
Odontomachus 22 44 Malaysia Goni et al., 1982
Odontomachus chelifer 44 4SM+40T Brazil Santos et al., 2010
Odontomachus haematodus 44 8SM+18ST+18A French Guiana Aguiar et al., 2020
Odontomachus hastatus 44 4SM+2ST+38T French Guiana Mariano et al., 2011
Odontomachus latidens 32 Indonesia Imai et al., 1985
Odontomachus latidens 15 Malaysia Imai et al., 1983
Odontomachus meinerti 44 4SM+6ST+34T Brazil Santos et al., 2010
Odontomachus rixosus 15 30 Malaysia Goni et al., 1982; Imai et al., 1983 B chomosomes 30, 30+1B
Odontomachus scalptus 44 2SM+16ST+26T French Guiana Mariano et al., 2011
Odontomachus simillimus 44 Sarawak Tjan et al., 1986
Odontomachus simillimus 22 44 Indonesia Imai et al., 1985
Odontomachus simillimus 22 44 Malaysia Goni et al., 1982; Imai et al., 1983
Odontoponera transversa 42 Indonesia Imai et al., 1985
Odontoponera transversa 46 14M+32A India Imai et al., 1984 the karyotype has been published only for the Indian poplations
Oecophylla longinoda 12 Crozier, 1970b
Oecophylla smaragdina 16 16M India Imai et al., 1984
Oecophylla smaragdina 8 Malaysia Crozier, 1970b
Ooceraea biroi 28 25M+3A India Imai et al., 1984 as ''Cerapachys biroi''
Opisthopsis rufithorax 50 Australia Imai et al., 1977
Orectognathus clarki 30 Australia Crozier, 1968d
Orectognathus darlingtoni 11 22 Australia Imai et al., 1977
Orectognathus versicolor 11 22 Australia Imai et al., 1977
Pachycondyla crassinoda 62 22M+40A Brazil Mariano et al., 2006c; Mariano et al., 2007
Pachycondyla harpax 90 16M+74A Brazil Velasco et al., 2014
Pachycondyla harpax 92 16M+76A Brazil Velasco et al., 2014
Pachycondyla harpax 96 12M+84A Brazil Mariano et al., 2006c; Mariano et al., 2007
Pachycondyla impressa 94 8M+86A Brazil Mariano et al., 2012
Pachycondyla striata 104 4M+100A Brazil Mariano et al., 2007
Papyrius nitidus 8 16 Australia Crozier, 1968a; Imai et al., 1977 as ''Iridomyrmex nitidus''
Paratrechina longicornis 16 Taiwan Hung et al., 1972
Paratrechina longicornis 16 Indonesia Imai et al., 1985
Paratrechina longicornis 8 16 14M+2A India Imai et al., 1984
Pheidole 16 Malaysia Goni et al., 1982; Imai et al., 1983
Pheidole 18 Australia Imai et al., 1977
Pheidole 18 Indonesia Imai et al., 1985
Pheidole 18 Malaysia Goni et al., 1982
Pheidole 18 Sarawak Tjan et al., 1986
Pheidole 18 18M India Imai et al., 1984
Pheidole 20 Australia Imai et al., 1977
Pheidole 20 Malaysia Goni et al., 1982
Pheidole 20 Malaysia Imai et al., 1983
Pheidole 20 20M India Imai et al., 1984
Pheidole 20 20M India Imai et al., 1984 near ''P. fossulata''
Pheidole 21 Malaysia Imai et al., 1983
Pheidole 22 Malaysia Imai et al., 1983
Pheidole 28 16M+12A India Imai et al., 1984 near ''P. watsoni''
Pheidole 30 18M+12A India Imai et al., 1984
Pheidole 32 Indonesia Imai et al., 1985
Pheidole 38 Malaysia Goni et al., 1982
Pheidole 38 Malaysia Imai et al., 1983
Pheidole 42 20M+22A India Imai et al., 1984 near ''P. grayi''
Pheidole 10 Japan Imai, 1969
Pheidole 10 20 Malaysia Goni et al., 1982; Imai et al., 1983
Pheidole 10 20 Malaysia Goni et al., 1982
Pheidole 10 20 Malaysia Imai et al., 1983
Pheidole 10 20 Sarawak Tjan et al., 1986
Pheidole 16 Malaysia Goni et al., 1982
Pheidole 17 Malaysia Goni et al., 1982
Pheidole 18 36 Malaysia Imai et al., 1983
Pheidole 9 Australia Crozier, 1966 'Pheidole concentrica'' group
Pheidole 9 18 Malaysia Imai et al., 1983
Pheidole 9 18 Sarawak Tjan et al., 1986
Pheidole binghamii 20 Indonesia Imai et al., 1985
Pheidole capellinii 20 Indonesia Imai et al., 1985
Pheidole dentata 20 USA Crozier, 1970b
Pheidole dentigula 10 20 USA Crozier, 1970b
Pheidole desertorum 20 20M USA Taber & Cokendolpher, 1988
Pheidole fallax 10 20 20M Uruguay Goni et al., 1983
Pheidole fervens 17 Crozier, 1975 as ''Pheidole noda''
Pheidole fervens 18 Crozier, 1975 as ''Pheidole noda''
Pheidole fervens 19 Crozier, 1975 as ''Pheidole noda''
Pheidole fervens 19 38 Japan Imai, 1969 as ''Pheidole noda''
Pheidole fervens 20 Crozier, 1975 as ''Pheidole noda''
Pheidole fervida 10 20 Japan Imai & Yosida, 1964; Imai, 1966; Imai, 1969
Pheidole hortensis 20 Indonesia Imai et al., 1985
Pheidole hyatti 20 20M USA Taber & Cokendolpher, 1988
Pheidole indica 20 20M India Imai et al., 1984 as ''Pheidole rotschana''
Pheidole latinoda 42 20M+22A India Imai et al., 1984
Pheidole megacephala 20 Malaysia Imai et al., 1983
Pheidole mus 12 12M India Imai et al., 1984
Pheidole nitidula 20 20M Uruguay Goni et al., 1983 as ''Pheidole strobeli''
Pheidole nodus 37 Japan Imai & Kubota, 1975
Pheidole nodus 39 Japan Imai & Kubota, 1975
Pheidole nodus 17 Japan Imai & Kubota, 1975
Pheidole nodus 18 Japan Imai & Kubota, 1975
Pheidole nodus 19 38 Japan Imai & Kubota, 1975
Pheidole nodus 20 Japan Imai & Kubota, 1975
Pheidole pallidula 24 Switzerland Hauschteck, 1961
Pheidole pallidula 10 20 Greece; Spain; Switzerland; Tunisia Hauschteck-Jungen & Jungen, 1983
Pheidole pallidula 10 20 8M+12SM Spain Palomeque et al., 1987; Palomeque et al., 1990b; Lorite et al., 2000
Pheidole plagiaria 20 Indonesia Imai et al., 1985
Pheidole porcula 20 20M USA Taber & Cokendolpher, 1988
Pheidole soritis 18 18M USA Taber & Cokendolpher, 1988 as ''Pheidole sitarches campestris''
Pheidole spininodis 20 20M Uruguay Goni et al., 1983
Pheidole subarmata 20 20M Uruguay Goni et al., 1983 as ''Pheidole cornutula''
Pheidole tepicana 18 18M USA Taber & Cokendolpher, 1988
Pheidole woodmasoni 18 18M India Imai et al., 1984
Philidris cordata 16 Malaysia Imai et al., 1983 as ''Iridomyrmex cordatus''
Philidris cordata 16 Sarawak Tjan et al., 1986 as ''Iridomyrmex cordatus''
Plagiolepis 18 Indonesia Imai et al., 1985
Plagiolepis 18 Malaysia Imai et al., 1983
Plagiolepis pygmaea 9 18 Croatia; Spain; Switzerland Hauschteck-Jungen & Jungen, 1983
Plagiolepis pygmaea 9 18 14M+2SM+2ST Spain Palomeque et al., 1993a
Plagiolepis schmitzii 9 18 Tunisia Hauschteck-Jungen & Jungen, 1983 as ''Plagiolepis barbara''
Plagiolepis schmitzii 9 18 12M+2A+2T Spain Palomeque et al., 1990b; Palomeque et al., 1993a as ''Plagiolepis barbara''
Platythyrea quadridenta 9 18 Malaysia Imai et al., 1983
Platythyrea sinuata 44 24M+20A Brazil Silva-Rocha et al., 2017
Platythyrea tricuspidata 92 Malaysia Imai et al., 1990
Platythyrea tricuspidata 93 Malaysia Imai et al., 1990
Platythyrea tricuspidata 94 Malaysia Imai et al., 1992
Platythyrea tricuspidata 96 Malaysia Imai et al., 1983 Corrected for 2n=92-94 in Imai 1990
Podomyrma adelaidae 49 Australia Imai et al., 1977
Podomyrma adelaidae 50 Australia Imai et al., 1977 B chromosome
Podomyrma adelaidae 51 Australia Imai et al., 1977 B chromosome
Pogonomyrmex apache 32 USA Taber et al., 1988
Pogonomyrmex badius 32 USA Taber et al., 1988
Pogonomyrmex barbatus 16 32 USA Taber et al., 1988
Pogonomyrmex brevispinosus 32 USA Taber et al., 1988
Pogonomyrmex californicus 32 USA Taber et al., 1988 as ''P. estebanius''
Pogonomyrmex comanche 16 32 USA Taber et al., 1988
Pogonomyrmex desertorum 32 USA Taber et al., 1988
Pogonomyrmex huachucanus 36 USA Taber et al., 1988
Pogonomyrmex imberbiculus 58 USA Taber et al., 1988
Pogonomyrmex imberbiculus 61 USA Taber et al., 1988 B chromosome
Pogonomyrmex imberbiculus 62 USA Taber et al., 1988 B chromosome
Pogonomyrmex imberbiculus 30 60 USA Taber et al., 1988 B chromosome
Pogonomyrmex magnacanthus 32 USA Taber et al., 1988
Pogonomyrmex maricopa 16 32 USA Taber et al., 1988
Pogonomyrmex montanus 32 USA Taber et al., 1988
Pogonomyrmex occidentalis 32 USA Mehlhop & Gardner, 1982; Taber et al., 1988
Pogonomyrmex rugosus 32 USA Taber et al., 1988
Pogonomyrmex subnitidus 16 32 USA Taber et al., 1988
Polyergus samurai 27 54 Japan Imai & Yosida, 1964; Imai, 1966; Imai, 1969
Polyrhachis 20 Malaysia Goni et al., 1982
Polyrhachis 42 Australia Imai et al., 1977
Polyrhachis 21 Malaysia Goni et al., 1982
Polyrhachis ammon 21 42 Australia Imai et al., 1977
Polyrhachis dives 21 1M+8SM+1ST+3A+8T Taiwan Hung et al., 1972
Polyrhachis gribodoi 48 Indonesia Imai et al., 1985
Polyrhachis hector 21 42 Malaysia Imai et al., 1983
Polyrhachis hippomanes 20 40 Japan Imai, 1969
Polyrhachis illaudata 14 28 Indonesia Imai et al., 1985
Polyrhachis illaudata 18 Malaysia Imai et al., 1983
Polyrhachis lacteipennis 42 6M+36A India Imai et al., 1984 as ''Polyrhachis simplex''
Polyrhachis lamellidens 42 Japan Imai, 1969
Polyrhachis rastellata 21 42 8M+34SM Malaysia Crozier, 1970b; Imai et al., 1983
Ponera 12 Indonesia Imai et al., 1985
Ponera 6 Japan Imai, 1969
Ponera japonica 12 Malaysia Imai et al., 1983
Ponera pennsylvanica 6 12 USA Hauschteck-Jungen & Jungen, 1983
Ponera scabra 7 Japan Imai et al., 1988a
Ponera scabra 8 Japan Imai et al., 1988a
Ponera scabra 3 Japan Crozier, 1975; Imai et al., 1988a
Ponera scabra 4 Japan Crozier, 1975; Imai et al., 1988a
Ponera scabra 4 8 2M+6SM Japan Imai & Kubota, 1972; Crozier, 1975
Prenolepis imparis 16 Switzerland Hauschteck, 1962
Prenolepis jerdoni 30 Malaysia Imai et al., 1983 B Chromosome
Prenolepis jerdoni 31 Malaysia Imai et al., 1983 B Chromosome
Prenolepis jerdoni 32 Malaysia Imai et al., 1983 B Chromosome
Prenolepis jerdoni 34 Malaysia Imai et al., 1983 B Chromosome
Prenolepis jerdoni 34 Malaysia Goni et al., 1982
Prenolepis jerdoni 36 Malaysia Goni et al., 1982
Prenolepis jerdoni 16 32 Malaysia Imai et al., 1983 B Chromosome
Prenolepis jerdoni 20 Malaysia Imai et al., 1983 B Chromosome
Prenolepis jerdoni 25 Malaysia Imai et al., 1983 B Chromosome
Prenolepis jerdoni 27 Malaysia Imai et al., 1983 B Chromosome
Pristomyrmex 22 Malaysia Goni et al., 1982
Pristomyrmex 14 Malaysia Imai et al., 1983
Pristomyrmex punctatus 12 24 Japan Imai & Yosida, 1964; Imai, 1966; Imai, 1969; Itow et al., 1984 as ''Pristomyrmex pungens''
Proatta butteli 32 Malaysia Goni et al., 1982
Probolomyrmex 28 Malaysia Goni et al., 1982
Proceratium 48 Sarawak Tjan et al., 1986
Proceratium silaceum 18 46 USA Crozier, 1970b
Prolasius 18 Australia Imai et al., 1977
Prolasius 9 18 Australia Imai et al., 1977
Pseudolasius 30 Indonesia Imai et al., 1985
Pseudolasius 30 Sarawak Tjan et al., 1986
Pseudolasius 14 Taiwan Hung et al., 1972 near ''Pseudolasius emeryi''
Pseudolasius 15 30 Malaysia Imai et al., 1983 B Chromosome polymorphism
Pseudolasius 17 30 Malaysia Imai et al., 1983 B Chromosome polymorphism
Pseudolasius 19 30 Malaysia Imai et al., 1983 B Chromosome polymorphism
Pseudolasius 8 Malaysia Imai et al., 1983
Pseudomyrmex 43 7M+36A Brazil Sposito et al., 2006 code PSW156
Pseudomyrmex 44 6M+38A Brazil Sposito et al., 2006
Pseudomyrmex 50 2M+48A Brazil Sposito et al., 2006 code PSW156
Pseudomyrmex gracilis 70 70A Brazil Sposito et al., 2006
Pseudomyrmex holmgreni 50 6M+44A or 2M+48A Brazil Sposito et al., 2006
Pseudomyrmex penetrator 24 22M+2A French Guiana Mariano et al., 2011
Pseudomyrmex penetrator 24 24M Brazil Sposito et al., 2006
Pseudomyrmex schuppi 24 5M+19A Brazil Sposito et al., 2006
Pseudoneoponera 60 Australia Imai et al., 1977
Pseudoneoponera rufipes 48 20M+28A India Imai et al., 1984 as ''Bothroponera rufipes''
Pseudoneoponera tridentata 28 Malaysia Imai et al., 1983 as ''Pachycondyla tridentata''
Pseudoponera gilberti 12 10M+2A Brazil Mariano et al., 2012 as ''Pachycondyla gilberti''
Pseudoponera gilberti 12 10M+2SM French Guiana Aguiar et al., 2020
Pseudoponera gilberti 14 12M+2AM Brazil Mariano et al., 2007 as ''Pachycondyla gilberti''
Pseudoponera gilberti 6 12 10M+2SM Brazil Correia et al., 2016
Pseudoponera stigma 12 12M Brazil Mariano et al., 2012 as ''Pachycondyla stigma''
Pseudoponera stigma 12 12M Brazil Mariano et al., 2007 as ''Pachycondyla stigma''
Pseudoponera stigma 14 14M Brazil Aguiar et al., 2020
Pseudoponera stigma 7 14 10M+4SM Brazil Correia et al., 2016
Pseudoponera succedanea 14 14M French Guiana Mariano et al., 2012 as ''Pachycondyla succedanea''
Recurvidris 24 18M+6A India Imai et al., 1984 as ''Trigonogaster'' sp.
Rhytidoponera 23 Australia Crozier et al., 1986
Rhytidoponera 46 Australia Crozier et al., 1986
Rhytidoponera 48 Australia Crozier et al., 1986
Rhytidoponera 49 Australia Crozier et al., 1986
Rhytidoponera 50 Australia Crozier et al., 1986
Rhytidoponera 52 Australia Crozier et al., 1986
Rhytidoponera aciculata 52 Australia Imai et al., 1977
Rhytidoponera chalybaea 42 Australia Imai et al., 1977
Rhytidoponera impressa 42 Australia Imai et al., 1977
Rhytidoponera lamellinodis 52 Australia Crozier et al., 1986
Rhytidoponera maniae 39 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera maniae 44 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera maniae 45 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera maniae 46 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera maniae 47 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera maniae 48 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera mayri 50 Australia Imai et al., 1977
Rhytidoponera metallica 22 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 23 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 24 Australia Crozier, 1969; Crozier et al., 1986 Robertsonian polymorphism
Rhytidoponera metallica 24 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 34 Australia Crozier et al., 1986 Robertsonian polymorphism
Rhytidoponera metallica 35 Australia Crozier et al., 1986 Robertsonian polymorphism
Rhytidoponera metallica 36 Australia Crozier et al., 1986 Robertsonian polymorphism
Rhytidoponera metallica 36 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 37 Australia Crozier et al., 1986 Robertsonian polymorphism
Rhytidoponera metallica 37 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 38 Australia Crozier et al., 1986 Robertsonian polymorphism
Rhytidoponera metallica 41 Australia Crozier, 1969 Robertsonian polymorphism
Rhytidoponera metallica 41 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 42 Australia Crozier, 1969 Robertsonian polymorphism
Rhytidoponera metallica 42 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 43 Australia Crozier, 1969; Crozier et al., 1986 Robertsonian polymorphism
Rhytidoponera metallica 43 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 44 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 46 Australia Crozier et al., 1986 Robertsonian polymorphism
Rhytidoponera metallica 46 Australia Imai et al., 1977 Robertsonian polymorphism
Rhytidoponera metallica 17 Australia Crozier, 1969 Robertsonian polymorphism
Rhytidoponera metallica 18 Australia Crozier, 1969 Robertsonian polymorphism
Rhytidoponera metallica 19 Australia Crozier, 1969 Robertsonian polymorphism
Rhytidoponera metallica 20 Australia Crozier, 1969 Robertsonian polymorphism
Rhytidoponera metallica 21 Australia Crozier, 1969 Robertsonian polymorphism
Rhytidoponera metallica 22 Australia Crozier, 1969 Robertsonian polymorphism
Rhytidoponera punctata 50 Australia Crozier et al., 1986
Rhytidoponera purpurea 38 Australia Imai et al., 1977
Rhytidoponera tasmaniensis 30 Australia Crozier et al., 1986
Rhytidoponera tasmaniensis 46 Australia Crozier et al., 1986
Rhytidoponera victoriae 21 42 Australia Crozier, 1969; Crozier, 1970b; Crozier et al., 1986; Imai et al., 1977
Sericomyrmex 25 50 44M+6SM Brazil Barros et al., 2013
Sericomyrmex amabilis 50 50M Panama Murakami et al., 1998
Solenopsis 38 Malaysia Goni et al., 1982 as ''Diplorhoptrum'' sp.
Solenopsis 11 Australia Crozier, 1966
Solenopsis aurea 16 32 USA Crozier, 1970b
Solenopsis fugax 22 Japan Imai, 1969
Solenopsis fugax 11 22 Switzerland Hauschteck, 1961
Solenopsis geminata 32 26M+6A India Imai et al., 1984
Solenopsis geminata 16 32 USA Crozier, 1970b
Solenopsis geminata 16 32 14M+12SM+6ST French Guiana Aguiar et al., 2020
Solenopsis invicta 16 32 USA Glancey et al., 1976
Solenopsis molesta 11 22 USA Crozier, 1970b
Solenopsis richteri 16 32 USA Glancey et al., 1976
Solenopsis saevissima 16 32 20M+10ST+2A Uruguay Goni et al., 1983
Solenopsis xyloni 16 32 28M+4A USA Taber & Cokendolpher, 1988
Stenamma brevicorne 4 8 USA Hauschteck, 1962; Hauschteck, 1963
Stenamma westwoodii 20 Crozier, 1975
Stigmacros 38 38A Australia Imai et al., 1977
Stigmacros 10 20 20M Australia Imai et al., 1977
Stigmatomma reclinatum 38 Malaysia Imai et al., 1983 as ''Amblyopone reclinata''
Strongylognathus huberi 14 28 Switzerland Hauschteck, 1962
Strumigenys 16 Malaysia Goni et al., 1982 as ''Smithistruma'' sp.
Strumigenys 38 Indonesia Imai et al., 1985 as ''Smithistruma'' sp.
Strumigenys 13 Crozier, 1975 as ''Smithistruma'' sp.
Strumigenys diabola 40 18SM+12ST+10A French Guiana Aguiar et al., 2020
Strumigenys dohertyi 12 24 Malaysia Imai et al., 1983 as ''Strumigenys taipingensis''
Strumigenys doriae 22 Indonesia Imai et al., 1985
Strumigenys friedae 24 Australia Imai et al., 1977
Strumigenys godeffroyi 40 Malaysia Imai et al., 1983
Strumigenys godeffroyi 44 Indonesia Imai et al., 1985
Strumigenys louisianae 4 4M Brazil AlvesSilva et al., 2014
Strumigenys mutica 36 Indonesia Imai et al., 1985 as ''Kyidris mutica''
Tapinoma 5 10 Malaysia Goni et al., 1982
Tapinoma erraticum 8 16 Switzerland Hauschteck-Jungen & Jungen, 1983
Tapinoma erraticum 8 16 12M+2SM+2ST Spain Palomeque et al., 1988; Lorite et al., 1998b
Tapinoma indicum 10 8M+2A India Imai et al., 1984
Tapinoma indicum 7 Crozier, 1975
Tapinoma melanocephalum 10 Indonesia Imai et al., 1985
Tapinoma melanocephalum 10 8M+2A India Imai et al., 1984
Tapinoma melanocephalum 5 10 USA Crozier, 1970a; Crozier, 1970b
Tapinoma nigerrimum 9 Tunisia Hauschteck-Jungen & Jungen, 1983
Tapinoma nigerrimum 9 18 10M+4SM+4ST Spain Palomeque et al., 1988; Palomeque et al., 1990a; Palomeque et al., 1990b; Palomeque et al., 1993a; Lorite et al., 1996a; Lorite et al., 1997; Lorite et al., 1998b; Lorite et al., 1999a; Lorite et al., 1999b; Lorite et al., 2002a
Tapinoma sessile 8 16 14M+2A USA Crozier, 1970a; Taber & Cokendolpher, 1988
Tapinoma simrothi 9 Crozier, 1975
Tapinoma simrothi 9 18 Spain; Tunisia Hauschteck-Jungen & Jungen, 1983
Technomyrmex 28 Indonesia Imai et al., 1985
Technomyrmex 28 8M+20A India Imai et al., 1984 'T. bicolor'' group
Technomyrmex 30 Indonesia Imai et al., 1985
Technomyrmex 30 Malaysia Goni et al., 1982; Imai et al., 1983
Technomyrmex albipes 16 Australia Imai et al., 1977
Technomyrmex albipes 16 16M India Imai et al., 1984
Technomyrmex albipes 9 Spain Lorite et al., 2012 as ''Tapinoma nigerum''
Technomyrmex albipes 9 18 16M+2A Australia Crozier, 1968a seven medium-size chromosomes metacentric and two small chromosomes one metacentric and one acrocentric.
Temnothorax 12 Greece Fischer, 1987 as ''Leptothorax'' cf. ''interruptus''
Temnothorax 14 France; Spain Fischer, 1987 as ''Leptothorax'' cf. ''lichtensteini''
Temnothorax 9 Croatia, France, Greece Fischer, 1987 as ''Leptothorax'' spp., ''tuberum'' group
Temnothorax adlerzi 10 Greece Douwes et al., 1988 as ''Epimyrma adlerzi''
Temnothorax affinis 9 France, Germany, Switzerland Fischer, 1987 as ''Leptothorax affinis''
Temnothorax albipennis 8 Germany Fischer, 1987 as ''Leptothorax albipennis'', ''L. tuberointerruptus'', ''Stenamma albipennis''
Temnothorax albipennis 8 Great Britain Orledge, 1998 as ''Leptothorax albipennis'', ''L. tuberointerruptus'', ''Stenamma albipennis''
Temnothorax algerianus 10 Douwes et al., 1988 as ''Epimyrma algeriana''
Temnothorax ambiguus 22 Canada Fischer, 1987 as ''Leptothorax ambiguus''
Temnothorax americanus 11 Canada; USA Fischer, 1987 as ''Harpagoxenus americanus''
Temnothorax andrei 10 USA Fischer, 1987 as ''Leptothorax andrei''
Temnothorax angustulus 23 France Fischer, 1987 as ''Leptothorax angustulus''
Temnothorax aveli 9 France Fischer, 1987 as ''Leptothorax rabaudi''
Temnothorax bernardi 10 Spain Buschinger et al., 1986; Fischer, 1987 as ''Epimyrma bernardi''
Temnothorax congruus 9 18 Japan Imai & Kubota, 1972 as ''Leptothorax congruus''
Temnothorax corsicus 10 France Buschinger et al., 1986; Fischer, 1987 as ''Epimyrma corsica''
Temnothorax corticalis 12 Germany Fischer, 1987 as ''Leptothorax corticalis''
Temnothorax corticalis 9 Crozier, 1975 as ''Leptothorax corticalis''
Temnothorax curvispinosus 23 USA Fischer, 1987 as ''Leptothorax curvispinosus''
Temnothorax duloticus 23 Canada Fischer, 1987 as ''Leptothorax duloticus''
Temnothorax duloticus 24 Canada Fischer, 1987 as ''Leptothorax duloticus''
Temnothorax duloticus 25 Canada Fischer, 1987 as ''Leptothorax duloticus''
Temnothorax exilis 11 Croatia; France; Italy; Turkey Fischer, 1987 as ''Leptothorax exilis''
Temnothorax exilis 17 Italy Fischer, 1987 as ''Leptothorax specularis''
Temnothorax flavicornis 11 Italy Fischer, 1987 as ''Leptothorax flavicornis''
Temnothorax gordiagini 10 Croatia Buschinger et al., 1983; Fischer, 1987 as ''Myrmoxenus gordiagini''
Temnothorax gredosi 17 Spain Fischer, 1987 as ''Leptothorax gredosi''
Temnothorax interruptus 24 Germany Hauschteck-Jungen & Jungen, 1983 as ''Leptothorax interruptus''
Temnothorax interruptus 12 Croatia; Germany Fischer, 1987 as ''Leptothorax interruptus''
Temnothorax kraussei 10 France, Italy, Spain Fischer, 1987 as ''Epimyrma kraussei''
Temnothorax kutteri 12 France Fischer, 1987 as ''Chalepoxenus kutteri''
Temnothorax lichtensteini 14 Croatia, France, Italy Fischer, 1987 as ''Leptothorax lichtensteini''
Temnothorax longispinosus 12 Canada Fischer, 1987 as ''Leptothorax longispinosus''
Temnothorax longispinosus 12 24 USA Crozier, 1970b as ''Leptothorax longispinosus''
Temnothorax luteus 21 France; Switzerland Fischer, 1987 as ''Leptothorax tristis''
Temnothorax mediterraneus 14 28 France, Italy, Spain Fischer, 1987 as ''Leptothorax kraussei''
Temnothorax melas 13 France Fischer, 1987 as ''Leptothorax melas''
Temnothorax muellerianus 12 Croatia; France; Greece; Italy Fischer, 1987 as ''Chalepoxenus siciliensis''
Temnothorax niger 18 France; Italy Fischer, 1987 as ''Leptothorax niger''
Temnothorax nigriceps 18 Switzerland Hauschteck, 1962 as ''Leptothorax nigriceps''
Temnothorax nigriceps 9 Croatia; France; Germany; Italy Fischer, 1987 as ''Leptothorax nigriceps''
Temnothorax nylanderi 11 Croatia, France, Germany Fischer, 1987 as ''Leptothorax nylanderi''
Temnothorax nylanderi 11 22 Switzerland Hauschteck-Jungen & Jungen, 1983 as ''Leptothorax nylanderi''
Temnothorax parvulus 14 France, Italy, Switzerland Fischer, 1987 as ''Leptothorax parvulus''
Temnothorax racovitzai 21 France Fischer, 1987 as ''Leptothorax racovitzai''
Temnothorax ravouxi 10 France; Germany; Italy; Switzerland Buschinger et al., 1986; Fischer, 1987 as ''Epimyrma ravouxi''
Temnothorax recedens 12 Italy Fischer, Karl, 1987 as ''Leptothorax recedens''
Temnothorax rottenbergii 11 Italy Fischer, 1987 as ''Leptothorax rottenbergii''
Temnothorax rougeti 8 Fischer, 1987 as ''Leptothorax rougeti'' (subspecies of ''Temnothorax unifasciatus'')
Temnothorax rugatulus 26 USA Taber & Cokendolpher, 1988 as ''Leptothorax rugatulus''
Temnothorax rugatulus 27 USA Taber & Cokendolpher, 1988 as ''Leptothorax rugatulus''
Temnothorax rugatulus 14 USA Fischer, 1987 as ''Leptothorax rugatulus''
Temnothorax schaumii 9 USA Hauschteck-Jungen & Jungen, 1983 as ''Leptothorax schaumii''
Temnothorax sordidulus 11 Croatia, Italy, Switzerland Fischer, 1987 as ''Leptothorax sordidulus''
Temnothorax spinosior 12 24 Japan Imai, 1966; Imai, 1969; Imai, 1974; Imai & Yosida, 1964; Fischer, 1987 as ''Leptothorax spinosior''
Temnothorax spinosus 16 Algeria Fischer, 1987 as ''Leptothorax spinosus''
Temnothorax stumperi 10 France; Switzerland Buschinger et al., 1986; Fischer, 1987 as ''Epimyrma stumperi''
Temnothorax tuberum 9 France, Sweden, Switzerland Fischer, 1987 as ''Leptothorax tuberum''
Temnothorax tuberum 9 18 Switzerland Hauschteck, 1962; Hauschteck-Jungen & Jungen, 1983 as ''Leptothorax tuberum''
Temnothorax unifasciatus 9 Croatia; France; Germany; Greece; Italy Fischer, 1987 as ''Leptothorax unifasciatus''
Temnothorax unifasciatus 9 18 Germany; Switzerland Hauschteck-Jungen & Jungen, 1983 as ''Leptothorax unifasciatus''
Tetramorium 14 Malaysia Goni et al., 1982
Tetramorium 18 Australia Imai et al., 1977 as ''Xiphomyrmex'' sp2.(ANIC)
Tetramorium 18 Australia Imai et al., 1977 as ''Xiphomyrmex'' sp4.(ANIC)
Tetramorium 18 Malaysia Goni et al., 1982 as ''Triglyphothrix'' sp. 2
Tetramorium 20 Australia Imai et al., 1977 as ''Xiphomyrmex'' sp3.(ANIC)
Tetramorium 20 Malaysia Goni et al., 1982 as ''Triglyphothrix'' sp. 1
Tetramorium 20 Sarawak Tjan et al., 1986
Tetramorium 20 Taiwan Hung et al., 1972 as ''Xiphomyrmex'' sp.
Tetramorium 22 Malaysia Goni et al., 1982
Tetramorium 24 Sarawak Tjan et al., 1986
Tetramorium 26 Malaysia Goni et al., 1982
Tetramorium 35 India Imai et al., 1984
Tetramorium 36 India Imai et al., 1984
Tetramorium adelphon 22 Indonesia Imai et al., 1985
Tetramorium brevidentatum 10 20 Indonesia Imai et al., 1985
Tetramorium caespitum 14 5SM+2ST+7A Spain Palomeque et al., 1987; Palomeque et al., 1990b; Lorite et al., 2000
Tetramorium caespitum 14 28 Japan Imai, 1966; Imai, 1969; Imai & Yosida, 1964
Tetramorium caespitum 14 28 Switzerland Hauschteck, 1961
Tetramorium eleates 28 Malaysia Imai et al., 1983
Tetramorium forte 14 4SM+3ST+7A Spain Palomeque et al., 1987; Lorite et al., 2000 as ''T. hispanicum''
Tetramorium guineense 22 Japan Imai, 1969
Tetramorium insolens 22 Indonesia Imai et al., 1985
Tetramorium kheperra 14 Indonesia Imai et al., 1985
Tetramorium lanuginosum 14 12M+2A India Imai et al., 1984 as ''Triglyphothrix lanuginosa''
Tetramorium pacificum 22 Indonesia Imai et al., 1985
Tetramorium pnyxis 20 Malaysia Imai et al., 1983
Tetramorium semilaeve 14 4SM+3ST+7A Spain Palomeque et al., 1987; Palomeque et al., 1990b; Lorite et al., 2000
Tetramorium seneb 20 Malaysia Imai et al., 1983
Tetramorium simillimum 14 India Imai et al., 1984
Tetramorium smithi 26 India Imai et al., 1984
Tetramorium smithi 26 Indonesia Imai et al., 1985
Tetramorium smithi 26 Sarawak Tjan et al., 1986
Tetramorium spinosum 13 26 26M USA Taber & Cokendolpher, 1988
Tetramorium walshi 14 14M India Imai et al., 1984 as ''Triglyphothrix walshi''
Tetraponera 42 Malaysia Imai et al., 1983
Tetraponera 44 Malaysia Goni et al., 1982
Tetraponera allaborans 16 Taiwan Hung et al., 1972
Trachymyrmex septentrionalis 10 20 20M Panama Murakami et al., 1998; Micolino et al., 2020
Trichomyrmex glaber 38 India Imai et al., 1984 near ''Monomorium glabrum'' current as ''Trichomyrmex glaber''
Trichomyrmex scabriceps 38 India Imai et al., 1984
Typhlomyrmex meire 10 20 Brazil Mariano et al., 2006b
Typhlomyrmex rogenhoferi 36 French Guiana Mariano et al., 2006b
Typhlomyrmex rogenhoferi 17 34 Brazil Mariano et al., 2006b
Typhlomyrmex rogenhoferi 19 38 Brazil Mariano et al., 2006b
Veromessor andrei 40 USA Taber & Cokendolpher, 1988
Vollenhovia 33 Malaysia Imai et al., 1983
Vollenhovia 34 Malaysia Imai et al., 1983
Vollenhovia 36 Indonesia Imai et al., 1985
Vollenhovia 40 Australia Imai et al., 1977
Vollenhovia 49 Malaysia Imai et al., 1983
Vollenhovia 50 Malaysia Imai et al., 1983
Vollenhovia 11 Crozier, 1975
Vollenhovia emeryi 18 36 Japan Imai & Yosida, 1964; Imai, 1966; Imai, 1969
Wasmannia auropunctata 32 16M+13SM+5ST French Guiana Aguiar et al., 2020
Wasmannia auropunctata 32 20M+12A Brazil deSouza et al., 2011
Zasphinctus steinheili 45 Australia Imai et al., 1977 as ''Sphinctomyrmex steinheili''
Zasphinctus steinheili 46 Australia Imai et al., 1977 as ''Sphinctomyrmex steinheili''

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