This species nests in open areas in soil or under stones, it may build unsightly nests of loose soil on lawns. Reproductives are found in the nest throughout the year, nuptial flights occur from May through September. Flights occur in the afternoon and are usually announced by large numbers of very aggressive ants milling around the nest entrance. These ants are aggressive with a painful sting, although not nearly as aggressive as the introduced red fire ant, Solenopsis invicta. They derive the name “native fire ant” from their sting. The species causes considerable damage to seed banks, kills newly hatched birds, girdles agricultural plants, attacks agricultural products, attacks electrical equipment, and is a serious, stinging kitchen pest. (Mackay and Mackay 2002)
- 1 Identification
- 2 Distribution
- 3 Biology
- 4 Castes
- 5 Nomenclature
- 6 References
- 7 References based on Global Ant Biodiversity Informatics
A member of the Solenopsis geminata species-group.
These ants are often concolorous dark in color, although they are also bicolored with a red head and mesosoma, and a black gaster. The gaster rarely has any light brown areas.
In Nevada this species is restricted to the Hot Desert region in the southern tip. Elevation range is from -160 ft. in Death Valley, California to 4500 ft. near Beatty, Nevada (Wheeler & Wheeler, 1986).
Distribution based on Regional Taxon Lists
Distribution based on AntMaps
Distribution based on AntWeb specimens
Check data from AntWeb
In Nevada, Wheeler & Wheeler (1986) examined 18 records. Five nests where under stones, 1 under rotten wood, 2 at the base of desert shrubs, 1 in chaff beside a nest of Pogonomyrmex rugosus, 1 in exposed soil with a crack as an entrance. One colony was described as populous and aggressive; the sting was annoying; seeds were found in the nest. Rissing (personal communication) reported that ants of this species raided colonies of Pheidole gilvescens, Pogonomyrmex californicus (numerous), Pogonomyrmex rugosus and Veromessor pergandei (numerous) on his study site in Boulder City, Nevada.
Hill (2007) provides the following notes on a raid by the army ant Neivamyrmex swainsonii:
On 27 June 2006, while on a collecting trip in west Texas, a colony of N. swainsonii was observed raiding a colony of Solenopsis xyloni. These observations were made just outside of Alpine, in Brewster County, Texas (30°20’46”N 103°41’39”W) at 1,548 m, behind a pavilion along a fencerow separating a hotel parking lot and a pasture. The activity occurred in an area measuring approximately 1x1.5 m that consisted of mostly bare soil and gravel with some forbs and Cynodon dactylon (L,) Pers (Poaceae) (Bermuda grass). The observations were made between 7:55 P.M.-9:10 P.M., and the temperature was 28.8°C.
While collections of ants were being made in the area, a large number of S. xyloni were observed, apparently relocating their colony from an old nest site to a new one approximately one meter away. Many N. swainsonii workers were emerging from three holes in the ground between these two locations, whereupon they attacked the S. xyloni workers and took their brood (eggs and pupa). In most cases, the S. xyloni workers only minimally defended their brood, before dropping it and running away. In other cases, the N. swainsonii took the brood from the mandibles of the S. xyloni workers after a brief skirmish. Several S. xyloni workers carrying brood apparently tried to evade the onslaught of their attackers by climbing onto a small forb. When a N. swainsonii worker ventured up the forb, the S. xyloni moved further up the plant until they were at the top. When the N. swainsonii neared them, the Solenopsis dropped their brood, and fell to the ground. The N. swainsonii workers also were observed also attacking male and female S. xyloni alates and carrying them underground after they were subdued. One S. xyloni worker also was observed being carried underground. Additionally, several N. swainsonii workers were seen entering and exiting the S. xyloni colony, but none of those exiting were observed carrying anything.
During the course of these observations, the movement trail of the Solenopsis became more obtuse as the Neivamyrmex pushed further into their ranks. A couple of workers of two other ant species, Pogonomyrmex rugosus and Novomessor cockerelli, were also moving throughout the area. The S. xyloni attacked both of these larger species when encountering them with seemingly greater aggressiveness than they exhibited for the more similar sized N. swainsonii, and in one case were able to kill one of the P. rugosus workers.
The observations ceased near sundown. The next morning the site was visited again, but there was no sign of either the Neivamyrmex or the Solenopsis. It would be interesting to know whether the Solenopsis were already moving their colony at the time and the Neivamyrmex took advantage of their vulnerability, or if the Solenopsis were moving because the Neivamyrmex already had attacked their original colony.
Association with Other Organisms
- This species is a host for the aphelinid wasp Aphytis melinus (a parasite) (Universal Chalcidoidea Database) (associate).
- This species is a host for the encyrtid wasp Comperiella bifasciata (a parasite) (Universal Chalcidoidea Database) (associate).
- This species is a host for the eucharitid wasp Orasema taii (a parasite) (Heraty, 1990; Chien & Heraty, 2018; Baker et al., 2019; Universal Chalcidoidea Database) (primary host; Solenopsis geminata X xyloni hybrid).
- This species is a host for the microsporidian fungus Kneallhazia solenopsae (Ascunce et al. 2010) (listed as S. geminata X S. xyloni hybrids).
The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.
- xyloni. Solenopsis xyloni McCook, 1879: 188, figs. 37, 38 (s.w.q.) U.S.A. Wheeler, W.M. 1915b: 396 (m.); Wheeler, G.C. & Wheeler, J. 1955c: 133 (l.); Taber & Cokendolpher, 1988: 95 (k.). Junior synonym of geminata: Mayr, 1886d: 460; Dalla Torre, 1893: 76. Revived from synonymy as subspecies of geminata: Wheeler, W.M. 1910g: 563; Wheeler, W.M. 1915b: 395. Revived status as species: Creighton, 1930b: 98; Creighton, 1950a: 232; Snelling, R.R. 1963: 9. Senior synonym of maniosa, pylades: Trager, 1991: 166.
- pylades. Solenopsis geminata r. pylades Forel, 1904d: 172 (q.) MEXICO. Forel, 1909a: 267 (w.); Forel, 1911c: 297 (m.); Bruch, 1916: 315 (m.). Raised to species: Forel, 1909a: 267; Bruch, 1914: 223; Forel, 1916: 459. Subspecies of saevissima: Santschi, 1923c: 267. Junior synonym of geminata: Emery, 1922e: 198; of saevissima: Wheeler, W.M. 1915b: 395; Wheeler, W.M. 1916g: 143; Wheeler, W.M. 1919c: 272; Creighton, 1930b: 70; of xyloni: Trager, 1991: 166.
- maniosa. Solenopsis geminata subsp. maniosa Wheeler, W.M. 1915b: 396 (w.q.m.) U.S.A. Subspecies of xyloni: Creighton, 1930b: 102. Junior synonym of xyloni: Creighton, 1950a: 232. Revived from synonymy and raised to species: Wheeler, G.C. & Wheeler, J. 1986g: 13. Junior synonym of xyloni: Trager, 1991: 166.
- n = 16, 2n = 32, karyotype = 28M+4A (USA) (Taber & Cokendolpher, 1988).
- Ajayi, O.S., Appel, A.G., Chen, L., Fadamiro, H.Y. 2020. Comparative Cutaneous Water Loss and Desiccation Tolerance of Four Solenopsis spp. (Hymenoptera: Formicidae) in the Southeastern United States. Insects 11, 418. (doi:10.3390/INSECTS11070418).
- Alvarez-Blanco, P., Broggi, J., Cerdá, X., González-Jarri, O., Angulo, E. 2020. Breeding consequences for a songbird nesting in Argentine ant’ invaded land. Biological Invasions 22, 2883–2898. (doi:10.1007/S10530-020-02297-3).
- Ascunce, M.S., Valles, S.M., Oi, D.H., Shoemaker, D., Plowes, R., Gilbert, L., LeBrun, E.G., Sanchez-Arroyo, H. & Sanchez-Peña, S. 2010. Molecular diversity of the microsporidium Kneallhazia solenopsae reveals an expanded host range among fire ants in North America. Journal of Invertebrate Pathology, 105(3), 279–288 (DOI 10.1016/j.jip.2010.07.008).
- Baker, A.J., Heraty, J.M., Mottern, J., Hang, J.Z., Hines, H.M., Lemmon, A.R., Lemmon, E.M. 2019. Inverse dispersal patterns in a group of ant parasitoids (Hymenoptera: Eucharitidae: Oraseminae) and their ant hosts. Systematic Entomology 45: 1–19 (doi:10.1111/syen.12371).
- Creighton, W. S. 1930b. The New World species of the genus Solenopsis (Hymenop. Formicidae). Proc. Am. Acad. Arts Sci. 66: 39-151 (page 98, revived status as species)
- Creighton, W. S. 1950a. The ants of North America. Bull. Mus. Comp. Zool. 104: 1-585 (page 232, revived status as species)
- Dalla Torre, K. W. von. 1893. Catalogus Hymenopterorum hucusque descriptorum systematicus et synonymicus. Vol. 7. Formicidae (Heterogyna). Leipzig: W. Engelmann, 289 pp. (page 76, Junior synonym of geminata)
- Hill, J.G. 2007. Observations of Neivamyrmex fallax Borgmier and Solenopsis xyloni McCook (Hymenoptera: Formicidae). Marginalia Insecta 2(2), 1-2.
- Hooper-Bùi, L.M., Rust, M.K. 1997. Food Preference and Patterns of Foraging Activity of the Southern Fire Ant, Solenopsis xyloni (McCook). Ann. Entomol. Soc. Amer. 90(2): 246-253.
- Hooper-Bùi, L.M., Rust, M.K. 1998. Factors that Induce Emergence and Flight of Solenopsis xyloni (McCook) Alates. Sociobiology 32(1): 63-75.
- Hooper-Bùi, L.M., Rust, M.K., Reierson, D.A. 2004. Predation of the Endangered California Least Tern, Sterna antillarum browni (Mearns) by the southern fire ant, Solenopsis xyloni (McCook). Sociobiology 43 (3): 401-418.
- Horna-Lowell, E., Neumann, K.M., O’Fallon, S., Rubio, A., Pinter-Wollman, N. 2021. Personality of ant colonies (Hymenoptera: Formicidae) – underlying mechanisms and ecological consequences. Myrmecological News 31: 47-59 (doi:10.25849/MYRMECOL.NEWS_031:047).
- Ipser, R.M., Brinkman, M.A., Gardner, W.A., Peeler, H.B. 2004. A survey of ground-dwelling ants (Hymenoptera: Formicidae) in Georgia. Florida Entomologist 87: 253-260.
- Mackay, W. P. and E. Mackay. 2002. The ants of New Mexico (Hymenoptera: Formicidae). Edwin Mellen Press, Lewiston, NY.
- McCook, H. C. 1880 . Formicariae. Pp. 182-189 in: Comstock, J. H. Report upon cotton insects. Washington, D.C.: Government Printing Office, 511 pp. (page 188, figs. 37, 38 soldier, worker, queen described)
- Mendonca, R., Melo, D., Prezoto, F. 2020. Evaluation of food selection of fire ant Solenopsis saevissima (Smith) (Hymenoptera: Formicidae) in resources with and without immature stages of Chrysomya albiceps (Wiedemann, 1819) (Diptera: Calliphoridae). Brazilian Journal of Development 6: 74119-74136.
- Moura, M.N., Cardoso, D.C., Cristiano, M.P. 2020. The tight genome size of ants: diversity and evolution under ancestral state reconstruction and base composition. Zoological Journal of the Linnean Society, zlaa135 (doi:10.1093/zoolinnean/zlaa135).
- Pitts, J.P., Camacho, G.P., Gotzek, D., McHugh, J.V., Ross, K.G. 2018. Revision of the fire ants of the Solenopisis saevissima species group (Hymenoptera: Formicidae). Proceedings of the Entomological Society of Washington, 120: 308–411.
- Ruano, F., Tinaut, A., Soler, J.J. 2000. High surface temperatures select for individual foraging in ants. Behavioral Ecology 11, 396-404.
- Sharaf, M.R., Gotzek, D., Guénard, B., Fisher, B.L., Aldawood, A.S., Al Dhafer, H.M., Mohamed, A.A. 2020. Molecular phylogenetic analysis and morphological reassessments of thief ants identify a new potential case of biological invasions. Scientific Reports 10, 12040 (doi:10.1038/s41598-020-69029-4).
- Snelling, R. R. 1963. The United States species of fire ants of the genus Solenopsis, subgenus Solenopsis Westwood, with synonymy of Solenopsis aurea Wheeler (Hymenoptera: Formicidae). Calif. Dep. Agric. Bur. Entomol. Occ. Pap. 3: 1-15 (page 9, revived status as species)
- Taber, S. W.; Cokendolpher, J. C. 1988. Karyotypes of a dozen ant species from the southwestern U.S.A. (Hymenoptera: Formicidae). Caryologia 41: 93-102 (page 95, karyotype described)
- Trager, J. C. 1991. A revision of the fire ants, Solenopsis geminata group (Hymenoptera: Formicidae: Myrmicinae). J. N. Y. Entomol. Soc. 99: 141-198 (page 166, Senior synonym of maniosa and pylades)
- Wheeler, G. C.; Wheeler, J. 1955c. The ant larvae of the myrmicine tribe Solenopsidini. Am. Midl. Nat. 54: 119-141 (page 133, larva described)
- Wheeler, W. M. 1910b. Ants: their structure, development and behavior. New York: Columbia University Press, xxv + 663 pp. (page 563, revived from synonymy as subspecies of geminata)
- Wheeler, W. M. 1915b. Some additions to the North American ant-fauna. Bull. Am. Mus. Nat. Hist. 34: 389-421 (page 396, male described; page 395, revived from synonymy as subspecies of geminata)
References based on Global Ant Biodiversity Informatics
- Adams T. A., W. J. Staubus, and W. M. Meyer. 2018. Fire impacts on ant assemblages in California sage scrub. Southwestern Entomologist 43(2): 323-334.
- Allred D. M. 1982. Ants of Utah. The Great Basin Naturalist 42: 415-511.
- Allred, D.M. 1982. The ants of Utah. Great Basin Naturalist 42:415-511.
- Author Unknown. 2004. Mirmecofauna (Hymenoptera: Formicidae) comu?n del estero El Salado y Puerto Vallarta, Jalisco, Me?xico. Dugesiana 11(1):13-20.
- Boulton A. M., Davies K. F. and Ward P. S. 2005. Species richness, abundance, and composition of ground-dwelling ants in northern California grasslands: role of plants, soil, and grazing. Environmental Entomology 34: 96-104
- Boulton A.M. and P.S. Ward. 2002. Ants. Chapter 5 in A New island Biogeography of the Sea of Cortes. T.J. Case, M.L. Cody and E. Ezcurra. Oxford university Press.
- Bruch C. 1914. Catálogo sistemático de los formícidos argentinos. Revista del Museo de La Plata 19: 211-234.
- Bruch C. 1916. Contribución al estudio de las hormigas de la provincia de San Luis. Revista del Museo de La Plata 23: 291-357.
- Chialvo P., D. A. Gotzek, D. Shoemaker, and K. G. Ross. 2018. Genetic analyses reveal cryptic diversity in the native North American fire ants (Hymenoptera: Formicidae: Solenopsis). Systematic Entomology 43: 109–122.
- Cokendolpher J.C., Reddell J.R., Taylor S.J, Krejca J.K., Suarez A.V. and Pekins C.E. 2009. Further ants (Hymenoptera: Formicidae) from caves of Texas [Hormigas (Hymenoptera: Formicdae) adicionales de cuevas de Texas]. Texas Memorial Museum Speleological Monographs, 7. Studies on the cave and endogean fauna of North America, V. Pp. 151-168
- Cole A. C., Jr. 1937. An annotated list of the ants of Arizona (Hymen.: Formicidae). [part]. Entomological News 48: 97-101.
- Cover S. P., and R. A. Johnson. 20011. Checklist of Arizona Ants. Downloaded on January 7th at http://www.asu.edu/clas/sirgtools/AZants-2011%20updatev2.pdf
- Crawley W. C. 1916. Ants from British Guiana. Ann. Mag. Nat. Hist. 8(17): 366-378.
- Creighton W. S. 1930. The New World species of the genus Solenopsis (Hymenop. Formicidae). Proceedings of the American Academy of Arts and Sciences 66: 39-151.
- Dash S. T. and L. M. Hooper-Bui. 2008. Species diversity of ants (Hymenoptera: Formicidae) in Louisiana. Conservation Biology and Biodiversity. 101: 1056-1066
- Dattilo W. et al. 2019. MEXICO ANTS: incidence and abundance along the Nearctic-Neotropical interface. Ecology https://doi.org/10.1002/ecy.2944
- Dean D. A., and S. R. Dean. 2018. A survey of the ant fauna and seasonal alate nuptial flights at two locations in South-Central Texas. Southwestern Entomologist 43(3): 639-647.
- Des Lauriers J., and D. Ikeda. 2017. The ants (Hymenoptera: Formicidae) of the San Gabriel Mountains of Southern California, USA with an annotated list. In: Reynolds R. E. (Ed.) Desert Studies Symposium. California State University Desert Studies Consortium, 342 pp. Pages 264-277.
- Deyrup M., C. Johnson, G. C. Wheeler, J. Wheeler. 1989. A preliminary list of the ants of Florida. Florida Entomologist 72: 91-101
- DuBois M. B. 1981. New records of ants in Kansas, III. State Biological Survey of Kansas. Technical Publications 10: 32-44
- DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-1344
- DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-1345
- DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-1346
- DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-1347
- DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-1348
- DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-1349
- DuBois M. B. 1985. Distribution of ants in Kansas: subfamilies Ponerinae, Ecitoninae, and Myrmicinae (Hymenoptera: Formicidae). Sociobiology 11: 153-1350
- Fisher B. L. 1997. A comparison of ant assemblages (Hymenoptera, Formicidae) on serpentine and non-serpentine soils in northern California. Insectes Sociaux 44: 23-33
- Forel A. 1911. Ameisen des Herrn Prof. v. Ihering aus Brasilien (Sao Paulo usw.) nebst einigen anderen aus Südamerika und Afrika (Hym.). Deutsche Entomologische Zeitschrift 1911: 285-312.
- Garcia-Martinez M. A., V. Vanoye-Eligio, O. R. Leyva-Ovalle, P. Zetina-Cordoba, M. J. Aguilar-Mendez, and M. Rosas-Mejia. 2019. Diversity of ants (Hymenoptera: Formicidae) in a sub-montane and sub-tropical cityscape of Northeastern Mexico. Sociobiology 66(3): 440-447.
- Greenberg L., M. Martinez, A. Tilzer, K. Nelson, S. Koening, and R. Cummings. 2015. Comparison of different protocols for control of the Red Imported Fire Ant, Solenopsis invicta Buren (Hymenoptera: Formicidae), in Orange County, California, including a list of co-occurring ants. Southwestern Entomologist 40(2): 297-305.
- Hess C. G. 1958. The ants of Dallas County, Texas, and their nesting sites; with particular reference to soil texture as an ecological factor. Field and Laboratory 26: 3-72.
- Hill J. G. 2007. Observations of Neivamyrmex fallax Borgmier and Solenopsis xyloni McCook (Hymenoptera: Formicidae). Marginalia Insecta 2: 1
- Holway D.A. 1998. Effect of Argentine ant invasions on ground-dwelling arthropods in northern California riparian woodlands. Oecologia. 116: 252-258
- Johnson R. Personnal Database. Accessed on February 5th 2014 at http://www.asu.edu/clas/sirgtools/resources.htm
- Johnson, R.A. and P.S. Ward. 2002. Biogeography and endemism of ants (Hymenoptera: Formicidae) in Baja California, Mexico: a first overview. Journal of Biogeography 29:10091026/
- Kaspari, M. and T.J. Valone. 2002. On Ectotherm Abundance in a Seasonal Environment-Studies of a Desert Ant Assemblage. Ecology 83(11):2991-2996
- La Rivers I. 1968. A first listing of the ants of Nevada. Biological Society of Nevada, Occasional Papers 17: 1-12.
- Luederwaldt H. 1918. Notas myrmecologicas. Rev. Mus. Paul. 10: 29-64.
- Luna, P., Y. Penaloza-Arellanes, A. L. Castillo-Meza, J. H. Garcia-Chavez, and W. Dattilo. 2018. Beta diversity of ant-plant interactions over day-night periods and plant physiognomies in a semiarid environment. Journal of Arid Environments 156: 69-76.
- MacGown, J.A and J.A. Forster. 2005. A preliminary list of the ants (Hymenoptera: Formicidae) of Alabama, U.S.A. Entomological News 116(2):61-74
- Mackay W. P., and E. E. Mackay. 2002. The ants of New Mexico (Hymenoptera: Formicidae). Lewiston, New York: Edwin Mellen Press, 400 pp.
- Mackay, W.P. and E. Mackay. XXXX. The Ants of New Mexico
- Mallis A. 1941. A list of the ants of California with notes on their habits and distribution. Bulletin of the Southern California Academy of Sciences 40: 61-100.
- Matsuda T., G. Turschak, C. Brehme, C. Rochester, M. Mitrovich, and R. Fisher. 2011. Effects of Large-Scale Wildfires on Ground Foraging Ants (Hymenoptera: Formicidae) in Southern California. Environmental Entomology 40(2): 204-216.
- McDonald D. L., D. R. Hoffpauir, and J. L. Cook. 2016. Survey yields seven new Texas county records and documents further spread of Red Imported Fire Ant, Solenopsis invicta Buren. Southwestern Entomologist, 41(4): 913-920.
- Michigan State University, The Albert J. Cook Arthropod Research Collection. Accessed on January 7th 2014 at http://www.arc.ent.msu.edu:8080/collection/index.jsp
- Miguelena J. G., and P. B. Baker. 2019. Effects of urbanization on the diversity, abundance, and composition of ant assemblages in an arid city. Environmental Entomology doi: 10.1093/ee/nvz069.
- Moody J. V., O. F. Francke, and F. W. Merickel. 1981. The distribution of fire ants, Solenopsis (Solenopsis) in Western Texas (Hymenoptera: Formicidae). Journal of the Kansas Entomological Society 54(3): 469-480.
- Moody J. V., and O. F. Francke. 1982. The Ants (Hymenoptera, Formicidae) of Western Texas Part 1: Subfamily Myrmicinae. Graduate Studies Texas Tech University 27: 80 pp.
- Nash M. S., W. G. Whitford, J. Van Zee, and K. M. Havstad. 2000. Ant (Hymenoptera: Formicidae) responses to environmental stressors in the Northern Chihuahuan Desert. Environ. Entomol, 29(2): 200-206.
- Ness, J.H., W.F. Morris and J.L. Bronstein. 2006. Integrating Quality and Quantity of Mutualistic Service to Contrast Ant Species Protecting Ferocactus Wislizeni. Ecology 87(4):912-921
- O'Keefe S. T., J. L. Cook, T. Dudek, D. F. Wunneburger, M. D. Guzman, R. N. Coulson, and S. B. Vinson. 2000. The Distribution of Texas Ants. The Southwestern Entomologist 22: 1-92.
- Reddell J. R., and J. C. Cokendolpher. 2001. Ants (Hymenoptera: Formicidae) from caves of Belize, Mexico, and California and Texas (U.S.A.) Texas. Texas Memorial Museum Speleological Monographs 5: 129-154.
- Rios-Casanova, L., A. Valiente-Banuet, and V. Rico-Gray. (2004). Las hormigas del Valle de Tehuacan (Hymenoptera: Formicidae): una comparacion con otras zonas aridas de Mexico. Acta Zoologica Mexicana 20: 37-54.
- Roeder K. A., and D. V. Roeder. 2016. A checklist and assemblage comparison of ants (Hymenoptera: Formicidae) from the Wichita Mountains Wildlife Refuge in Oklahoma. Check List 12(4): 1935.
- Ríos-Casanova, L., A. Valiente-Banuet and V. Rico-Gray. 2004. Las hormigas del Valle de Tehuacan (Hymenoptera: Formicidae): Una comparacion con otras zonas aridas de Mexico. Acta Zoologica Mexicana 20(1):37-54
- Smith M. R. 1934. A list of the ants of South Carolina. Journal of the New York Entomological Society 42: 353-361.
- Smith M. R. 1935. A list of the ants of Oklahoma (Hymen.: Formicidae). Entomological News 46: 235-241.
- Smith M. R. 1936. A list of the ants of Texas. Journal of the New York Entomological Society 44: 155-170.
- Staubus W. J., E. S. Boyd, T. A. Adams, D. M. Spear, M. M. Dipman, W. M. Meyer III. 2015. Ant communities in native sage scrub, non-native grassland, and suburban habitats in Los Angeles County, USA: conservation implications. Journal of Insect Conservervation 19:669–680
- Taber S. W., and J. C. Cokendolpher. 1988. Karyotypes of a dozen ant species from the southwestern U.S.A. (Hymenoptera: Formicidae). Caryologia 41: 93-102.
- Van Pelt, A. 1983. Ants of the Chisos Mountains, Texas (Hymenoptera: Formicidae) . Southwestern Naturalist 28:137-142.
- Varela-Hernandez, F., M. Rocha-Ortega, R. W. Jones, and W. P. Mackay. 2016. Insectos: Hormigas (Formicidae) del estado de Queretaro, Mexico. Pages 397-404 in W. Jones., and V. Serrano-Cardenas, editors. Historia Natural de Queretaro. Universidad Autonoma de Queretaro, Mexico.
- Vasquez-Bolanos M. 2011. Checklist of the ants (Hymenoptera: Formicidae) from Mexico. Dugesiana 18(1): 95-133.
- Vásquez-Bolaños M. 2011. Lista de especies de hormigas (Hymenoptera: Formicidae) para México. Dugesiana 18: 95-133
- Ward P. S. 1987. Distribution of the introduced Argentine ant (Iridomyrmex humilis) in natural habitats of the lower Sacramento Valley and its effects on the indigenous ant fauna. Hilgardia 55: 1-16
- Ward P.S. and A.M. Boulton. 2002. Checklist of the ants of the Gulf of California Islands. In Island Biogeography of the sea of Cortes. T.J. Case, M.L. Cody and E. Ezcurra Editors. 690 pp.
- Warren, L.O. and E.P. Rouse. 1969. The Ants of Arkansas. Bulletin of the Agricultural Experiment Station 742:1-67
- Wheeler G. C., and J. Wheeler J. 1989. A checklist of the ants of Oklahoma. Prairie Naturalist 21: 203-210.
- Wheeler G. C., and J. Wheeler. 1986. The ants of Nevada. Los Angeles: Natural History Museum of Los Angeles County, vii + 138 pp.
- Wheeler W. M. 1905. The ants of the Bahamas, with a list of the known West Indian species. Bulletin of the American Museum of Natural History 21: 79-135.
- Wheeler W. M. 1915. Some additions to the North American ant-fauna. Bull. Am. Mus. Nat. Hist. 34: 389-421.
- Wheeler, G.C. and J. Wheeler. 1985. A checklist of Texas ants. Prairie Naturalist 17:49-64.
- Whitford W. G. 1978. Structure and seasonal activity of Chihuahua desert ant communities. Insectes Sociaux 25(1): 79-88.
- Whitford W. G., J. Van Zee, M. S. Nash, W. E. Smth, and J. E. Herrick. 1999. Ants as indicators of exposure to environmental stressors in North American desert grasslands. Environmental Monitoring and Assessment 54: 143171.
- Wolkovich, E.M., D.T. Bolger and D.A. Holway. 2009. Complex responses to invasive grass litter by ground arthropods in a Mediterranean scrub ecosystem. Oecologia 161(4): 697-708
- Young J., and D. E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publication. Oklahoma Agricultural Experimental Station 71: 1-42.
- Young, J. and D.E. Howell. 1964. Ants of Oklahoma. Miscellaneous Publications of Oklahoma State University MP-71