Workers were collected from Monterey County (CA) on Eriogonum.
Bolton (2000) - A member of the Strumigenys rostrata-group. Within the group the dentition and adiastemate condition of the mandible allies this species with Strumigenys arizonica and Strumigenys rostrata (see there). The apical sculpture of the gastral segments is unique in the group and immediately identifies californica.
Keys including this Species
Distribution based on Regional Taxon Lists
Distribution based on AntMaps
Distribution based on AntWeb specimens
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The following information is derived from Barry Bolton's New General Catalogue, a catalogue of the world's ants.
- californica. Smithistruma (Smithistruma) californica Brown, 1950b: 40 (q.) U.S.A. Ward, 1988: 120 (w.). Combination in Pyramica: Bolton, 1999: 1673; in Strumigenys: Baroni Urbani & De Andrade, 2007: 116. See also: Brown, 1953g: 85; Bolton, 2000: 126.
Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.
Bolton (2000) - In the original description of the holotype queen Brown (1950b) mentions that the head has one or two transverse rows of elongate erect hairs near the occipital margin. Of the worker Ward (1988) says that the head has no sign of flagellate or linear hairs, and this also applies to workers from San Lucas. The LACM paratype queen, as well as having a few standing hairs near the occipital margin, has a short simple apicoscrobal curved hair, this latter hair not being recorded by either Brown or Ward. It is possible that these differences may reflect caste-specific characters, or that the workers may have had their longer cephalic hairs abraded. But it may imply that two sibling species are present. Material of californica is so sparse that no conclusion can be reached at present.
Bolton (2000) - HL 0.60-0.62, HW 0.38, CI 62, MI 16, AL 0.61 (measurements from Ward, 1988). Basal lamella of mandible followed without a diastema by the principal row of 5 acute teeth. First gastral tergite with sharply defined basigastral costulae that extend about one-quarter the length of the sclerite. Behind this the tergite smooth until the apical quarter or so, where there is a distinct transverse band of fine reticulate and longitudinally striate sculpture. Similar sculpture present at apex of first gastral sternite and at apices of tergites 2 and 3. Pronotal humeral hair long fine and flagellate.
Holotype. Total length (TL) 2.41, head length (HL) 0.64, cephalic index (CI) 67, mandibulo-cephalic index (MI) 17. Closely related to Sm. rostrata (Emery) and to the Japanese species rostrataeformis Brown and incerta Brown. Differs from rostrata in considerably smaller size and in the shorter, broader hairs of the free clypeal border, particularly those on the anterolateral "corners" of the clypeus. Humeral angles each with a distinctly flagelliform hair; gastric hairs flagelliform, erect, distinctly more numerous than are those of either worker or female of rostrata. Propodeal teeth short, compressed, subtended by convexly expanded infradental lamellae. Dorsum of petiole with sculpture more or less effaced, subopaque to weakly shining in varying lights. Mandibles shorter than those of rostrata, but appearing similarly toothed; no diastema evident, even in partially open mandible. Differs from the types of incerta (female and worker) and rostrataeformis in details of pilosity and in proportions of head, clypeus and mandibles. The anterior median clypeal emargination is present but very weak, not so well developed as in incerta, but perhaps a little stronger than in the majority of rostrata females. Color ferrugineous yellow; apex of gaster slightly darkened. Remnants of wings present.
Holotype and single paratype (see below) taken at Claremont, California (C. F. Baker). The data go no farther, as might be wished in the case of the only certain record for a member of the genus from west of the Great Plains in North America. Whether this locality is in the irrigated, intensively cultivated region around Claremont or in one of the canyons near the city may make the difference between an ant introduced from the Orient or one native to California. The paratype, also a female with large parts of the wings remaining, unfortunately lacks a head. The two females were probably captured during or just after the nuptial flight. These two (holotype and paratype) specimens, now in the Museum of Comparative Zoology, are the basis for the record of rostrata from California. Wheeler made the original determination, and this record appeared in Smith's revision of 1931. Dr. W. S. Creighton has rightly called this record to question in his book on North American ants (1950).
- Baroni Urbani, C. & De Andrade, M.L. 2007. The ant tribe Dacetini: limits and constituent genera, with descriptions of new species. Annali del Museo Civico di Storia Naturale “G. Doria” 99:1-191.
- Bolton, B. 1999. Ant genera of the tribe Dacetonini (Hymenoptera: Formicidae). Journal of Natural History. 33:1639-1689. PDF (page 1673, Combination in Pyramica)
- Bolton, B. 2000. The ant tribe Dacetini. Memoirs of the American Entomological Institute. 65:1-1028. (page 126, catalogue)
- Brown, W. L., Jr. 1950d. Preliminary descriptions of seven new species of the dacetine ant genus Smithistruma Brown. Trans. Am. Entomol. Soc. 76: 37-45 PDF (page 40, queen described)
- Brown, W. L., Jr. 1953. Revisionary studies in the ant tribe Dacetini. American Midland Naturalist. 50:1-137. PDF (page 85, see also)
- Ward, P. S. 1988. Mesic elements in the western Nearctic ant fauna: taxonomic and biological notes on Amblyopone, Proceratium, and Smithistruma (Hymenoptera: Formicidae). J. Kans. Entomol. Soc. 61: 102-124 (page 120, worker described)