Tetramorium bicarinatum

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Tetramorium bicarinatum
Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Hymenoptera
Family: Formicidae
Subfamily: Myrmicinae
Tribe: Crematogastrini
Genus: Tetramorium
Species group: bicarinatum
Species: T. bicarinatum
Binomial name
Tetramorium bicarinatum
(Nylander, 1846)

Tetramorium bicarinatum casent0005826 profile 1.jpg

Tetramorium bicarinatum casent0005826 dorsal 1.jpg

Specimen labels

Synonyms


Common Name
Oo-shiwa-ari
Language: Japanese
Guinea Ant
Language: English

Tetramorium bicarinatum is one of the most successful, abundant, and widespread tramp species within the genus Tetramorium. It occurs in almost all tropical and subtropical habitats around the world, except for most of the Afrotropical, and also has been transferred to temperate countries where it can survive in constantly heated buildings (botanical gardens, zoos, etc.). The species is known to occur in most of the Oriental and Indo-Australian regions, including almost all Pacific island systems, the New World, Europe, the Malagasy region, and on several islands in the Atlantic and Indian Oceans (Hita Garcia and Fisher 2011). It is found in grassland and areas of bare coastal soil in Japan, where it prefers more open and dryer situations than Tetramorium nipponense. Nests are found under stones and logs. (Japanese Ant Image Database)

At a Glance • Highly invasive  • Supercolonies  

Identification

Hita Garcia and Fisher (2011) - A member of the T. bicarinatum species complex in the Tetramorium bicarinatum species group. Tetramorium bicarinatum can be well identified within the T. bicarinatum group in the Malagasy region because of its distinctly sculptured mandibles, rectangular nodiform petiolar node with antero- and posterodorsal angles at about the same height, and its characteristic bicolouration.

Keys including this Species

Distribution

Worldwide throughout tropical and subtropical regions (widespread tramp species). It has been reported from the Mediterranean in the Iberian Peninsula (Reyes & Espadaler, 2005), Italy (Limonta et al., 2003), Montenegro (Petrov, 2008) and Morocco (Taheri et al., 2017) (see Gomez, 2017).

Latitudinal Distribution Pattern

Latitudinal Range: 29.281° to -10.476379°.

   
North
Temperate
North
Subtropical
Tropical South
Subtropical
South
Temperate

Distribution based on Regional Taxon Lists

Afrotropical Region: Cape Verde, Comoros, Eritrea, Nigeria, United Arab Emirates, Yemen.
Australasian Region: Australia, Lord Howe Island, New Caledonia, New Zealand, Norfolk Island.
Indo-Australian Region: American Samoa, Borneo, Fiji, Guam, Hawaii, Indonesia, Kiribati, Krakatau Islands, Malaysia, Marshall Islands, Micronesia (Federated States of), New Guinea, Niue, Philippines, Samoa, Singapore, Solomon Islands, Tokelau, Tonga, Vanuatu.
Malagasy Region: Madagascar, Mayotte, Seychelles.
Nearctic Region: United States (type locality).
Neotropical Region: Aruba, Barbados, Costa Rica, Cuba, Dominican Republic, Ecuador, French Guiana, Galapagos Islands, Greater Antilles, Grenada, Guadeloupe, Guyana, Haiti, Honduras, Mexico, Netherlands Antilles, Panama, Peru, Puerto Rico, Trinidad and Tobago.
Oriental Region: Bangladesh, Bhutan, India, Nepal, Nicobar Island, Sri Lanka, Thailand, Vietnam.
Palaearctic Region: Austria, Belgium, Canary Islands, China, Cyprus, Denmark, Germany, Hungary, Iberian Peninsula, Israel, Italy, Japan, Malta, Montenegro, Republic of Korea, Spain, Sweden, Türkiye.

Hita Garcia and Fisher (2011) - Its native range was originally thought to be in Africa by a number of authors (Creighton, 1950; Brown, 1957; Taylor & Wilson, 1961; Wilson & Taylor, 1967). Yet the species is not even native to the Afrotropical region, and is only known there from very few introductions (Bolton, 1979, 1980). It seems that the Afrotropical members of the T. bicarinatum group, all endemic to this region, have the ability to hinder the establishment of T. bicarinatum. Outside of Africa, in the Malagasy, Neotropical, and parts of the Palaearctic and Neartic regions, these T. bicarinatum group species are absent, and T. bicarinatum can set up stable populations. The situation is different in the Oriental and Indo-Australian regions because T. bicarinatum is widespread and common but the T. bicarinatum species group is also present with a number of species closely related to T. bicarinatum. This fact led Bolton (1977, 1979, 1980) to the conclusion that the native range of T. bicarinatum is most probably in South East Asia, from which it was anthropogenically transferred to the rest of its actual, immense distribution range. This view is supported by several authors (McGlynn, 1999; Deyrup, 2000; Astruc et al., 2001) and we concur with this opinion.

Distribution based on AntMaps

AntMapLegend.png

Distribution based on AntWeb specimens

Check data from AntWeb

Countries Occupied

Number of countries occupied by this species based on AntWiki Regional Taxon Lists. In general, fewer countries occupied indicates a narrower range, while more countries indicates a more widespread species.
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Estimated Abundance

Relative abundance based on number of AntMaps records per species (this species within the purple bar). Fewer records (to the left) indicates a less abundant/encountered species while more records (to the right) indicates more abundant/encountered species.
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Biology

Regional Notes

Australia

Heterick (2009) - Common in Perth gardens and with a wide distribution throughout Australia. The range for this species given by Brown and Taylor (1985) is probably understated; e.g. I have seen T . bicarinatum in Port Augusta, SA, although this State is not listed by the former authors. Unlike some other exotic myrmicines in Australia, this species does not seem to adversely affect the native ant fauna.

Malagasy region

Hita Garcia and Fisher (2011) - T. bicarinatum is almost surely not native to the Malagasy region and can be considered introduced. It is now a common species well-established in most of Eastern and Northern Madagascar, and also can be found in some tropical dry forest or gallery forest habitats in the western part of the country. It seems completely absent from the more arid Southwest and the High Plateau. In addition, T. bicarinatum can be commonly encountered on most of the surrounding islands, e.g. the Comoros, Mayotte, Seychelles, Reunion, and Mauritius.

Florida USA

A moderately common species occurring as far north and west as Walton county. Nests are usually in dead wood or hollow stems or under loose bark, usually at the edges of wet areas. This species is not particularly associated with disturbed habitats, and if it were not a notorious tropical tramp (Bolton 1979), one would never suspect from its ecology that this was an introduced species. Pest status: minor; known to occasionally enter houses, and is capable of stinging. First published Florida record: Emery 1895. (Deyrup, Davis & Cover, 2000.)

Association with Other Organisms

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  • This species is a mutualist for the aphid Aphis gossypii (a trophobiont) (Idechiil et al., 2007; Saddiqui et al., 2019).
  • This species is a mutualist for the aphid Pentalonia nigronervosa (a trophobiont) (Idechiil et al., 2007; Saddiqui et al., 2019).

Flight Period

X X
Jan Feb Mar Apr May Jun Jul Aug Sep Oct Nov Dec

Source: antkeeping.info.

Castes

Worker

Additional images can be found on the species category page.

Images from AntWeb

Tetramorium bicarinatum casent0059100 head 1.jpgTetramorium bicarinatum casent0059100 profile 1.jpgTetramorium bicarinatum casent0059100 dorsal 1.jpgTetramorium bicarinatum casent0059100 label 1.jpg
Worker. Specimen code casent0059100. Photographer Erin Prado, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Tetramorium bicarinatum casent0102404 head 1.jpgTetramorium bicarinatum casent0102404 profile 1.jpgTetramorium bicarinatum casent0102404 dorsal 1.jpgTetramorium bicarinatum casent0102404 label 1.jpg
Worker. Specimen code casent0102404. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by NHMUK, London, UK.
Tetramorium bicarinatum casent0102737 head 1.jpgTetramorium bicarinatum casent0102737 profile 1.jpgTetramorium bicarinatum casent0102737 dorsal 1.jpgTetramorium bicarinatum casent0102737 label 1.jpg
Worker. Specimen code casent0102737. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by ABS, Lake Placid, FL, USA.
Tetramorium bicarinatum casent0125127 head 1.jpgTetramorium bicarinatum casent0125127 profile 1.jpgTetramorium bicarinatum casent0125127 dorsal 1.jpgTetramorium bicarinatum casent0125127 label 1.jpg
Worker. Specimen code casent0125127. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.

Queen

Images from AntWeb

Tetramorium bicarinatum casent0135255 head 1.jpgTetramorium bicarinatum casent0135255 profile 1.jpgTetramorium bicarinatum casent0135255 dorsal 1.jpgTetramorium bicarinatum casent0135255 label 1.jpg
Queen (alate/dealate). Specimen code casent0135255. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Tetramorium bicarinatum casent0064305 head 1.jpgTetramorium bicarinatum casent0064305 profile 1.jpgTetramorium bicarinatum casent0064305 profile 2.jpgTetramorium bicarinatum casent0064305 dorsal 1.jpgTetramorium bicarinatum casent0064305 label 1.jpg
Queen (alate/dealate). Specimen code casent0064305. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.

Male

Images from AntWeb

Tetramorium bicarinatum casent0064306 head 1.jpgTetramorium bicarinatum casent0064306 profile 1.jpgTetramorium bicarinatum casent0064306 profile 2.jpgTetramorium bicarinatum casent0064306 dorsal 1.jpgTetramorium bicarinatum casent0064306 label 1.jpg
Male (alate). Specimen code casent0064306. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CAS, San Francisco, CA, USA.
Tetramorium bicarinatum casent0102736 head 1.jpgTetramorium bicarinatum casent0102736 profile 1.jpgTetramorium bicarinatum casent0102736 profile 2.jpgTetramorium bicarinatum casent0102736 dorsal 1.jpgTetramorium bicarinatum casent0102736 label 1.jpg
Male (alate). Specimen code casent0102736. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by ABS, Lake Placid, FL, USA.
Tetramorium bicarinatum casent0173283 head 1.jpgTetramorium bicarinatum casent0173283 profile 1.jpgTetramorium bicarinatum casent0173283 profile 2.jpgTetramorium bicarinatum casent0173283 profile 3.jpgTetramorium bicarinatum casent0173283 dorsal 1.jpgTetramorium bicarinatum casent0173283 label 1.jpg
Male (alate). Specimen code casent0173283. Photographer April Nobile, uploaded by California Academy of Sciences. Owned by CDRS, Galapagos, Ecuador.

Nomenclature

The following information is derived from Barry Bolton's Online Catalogue of the Ants of the World.

  • bicarinatum. Myrmica bicarinata Nylander, 1846b: 1061 (w.q.) U.S.A. Forel, 1891b: 151 (m., misidentified as guineense); Wheeler, G.C. & Wheeler, J. 1954d: 449 (l., misidentified as guineense); Wheeler, W.M. 1924c: 136 (gynandromorph, misidentified as guineense). Combination in Tetramorium: Mayr, 1862: 740; Bolton, 1977: 94. Junior synonym of Formica guineensis Fabricius, 1793: 357 (now in Pheidole): Mayr, 1862: 740. Revived from synonymy and senior synonym of cariniceps (and its junior synonym kollari), modesta Smith, reticulata: Bolton, 1977: 94. [Note. The names cariniceps, kollari and reticulata had previously been incorrectly synonymised with guineensis Fabricius by Roger, 1862c: 293; modesta Smith was wrongly synonymised with guineensis by Donisthorpe, 1932c: 463.] See also: Bolton, 1979: 164; Hita Garcia & Fisher, 2011: 18.
  • cariniceps. Myrmica cariniceps Guérin-Méneville, 1852: 79 (w.) DOMINICA. Combination in Tetramorium: Roger, 1861b: 171. Senior synonym of kollari: Roger, 1861b: 171. Junior synonym of guineense: Roger, 1862c: 293; of bicarinatum: Bolton, 1977: 94.
  • kollari. Myrmica kollari Mayr, 1853c: 283 (w.q.m.) AUSTRIA. Junior synonym of cariniceps: Roger, 1861b: 171.
  • modesta. Myrmica modesta Smith, F. 1860b: 108 (w.) INDONESIA (Batjan I.). [Unresolved junior primary homonym of modesta Foerster, above.] Combination in Tetramorium: Donisthorpe, 1932c: 463. Junior synonym of guineense: Donisthorpe, 1932c: 463; of bicarinatum: Bolton, 1977: 94.
  • reticulata. Myrmica reticulata Smith, F. 1862b: 33 (w.) PANAMA. Combination in Tetramorium: Mayr, 1862: 740. Junior synonym of guineense: Roger, 1862c: 293; of bicarinatum: Bolton, 1977: 94.

The following notes on F. Smith type specimens have been provided by Barry Bolton (details):

Myrmica modesta

Three worker syntypes (2 on one card, 1 on another) in Oxford University Museum of Natural History. Labelled “Bac 19.”

Two other specimens, labelled “Gil. 13,” are also present.

Unless otherwise noted the text for the remainder of this section is reported from the publication that includes the original description.

Type Material

  • Myrmica bicarinata: Syntype, worker(s), queen(s) (apparently lost), California, United States.

Description

Worker

Hita Garcia and Fisher (2011 ) - HL 0.760–0.910 (0.848); HW 0.655–0.810 (0.740); SL 0.510–0.625 (0.575); EL 0.185–0.220 (0.201); PW 0.480–0.580 (0.532); WL 0.940–1.120 (1.037); PSL 0.175–0.240 (0.208); PTL 0.230–0.290 (0.262); PTH 0.260–0.330 (0.299); PTW 0.235–0.300 (0.264); PPL 0.220–0.280 (0.247); PPH 0.250–0.315 (0.292); PPW 0.295–0.365 (0.329); CI 85–89 (87); SI 75–80 (78); OI 26–28 (27); PSLI 22–26 (24); PeNI 46–52 (50); LPeI 84–93 (88); DPeI 96–106 (101); PpNI 59–66 (62); LPpI 81–91 (85); DPpI 128–139 (133); PPI 120–132 (124) (23 measured).

Head distinctly longer than wide (CI 85–89). Anterior clypeal margin with distinct median impression. Frontal carinae strongly developed, ending shortly before posterior head margin. Antennal scrobes weakly developed, narrow, shallow, and posterior and ventral margins never differentiated, as long as frontal carinae. Antennal scapes of moderate length, not reaching posterior margin of head (SI 75–80). Eyes relatively large (OI 26–28), with 11 to 14 ommatidia in longest row. Mesosomal outline dorsally flat, metanotal groove absent. Propodeal spines medium-sized to long (PSLI 22–26). Propodeal lobes small, triangular, and acute. Petiolar node rectangular nodiform, anterior and posterior faces roughly parallel, anterodorsal and posterodorsal angles at about same height, in dorsal view weakly longer than wide to weakly wider than long (DPeI 96–106), in lateral view higher than long (LPeI 84–93). Postpetiole in profile roughly rounded and higher than long (LPpI 81–91), in dorsal view distinctly wider than long (DPpI 128–139). Postpetiole in profile less voluminous than petiolar node, in dorsal view always distinctly wider than petiole (PPI 120–132). Mandibles distinctly longitudinally striate. Clypeus longitudinally rugose, usually with three strong rugae. Most of head with reticulate-rugose sculpturation, area between frontal carinae from posterior clypeal margin to posterior eye margin irregularly rugose with few cross-meshes; ground sculpturation on head weak and faint. Mesosoma and waist segments reticulate-rugose without ground sculpture. Gaster with basigastral costulae on the first tergite, remainder completely unsculptured, smooth, and shiny. All dorsal surfaces of head, mesosoma, waist segments, and gaster with numerous long, standing hairs; hairs on antennal scapes and tibiae appressed to suberect. Head, mesosoma, waist segments, and legs yellowish orange to orange brown, gaster always much darker, usually dark brown to blackish brown.

Type Material

Hita Garcia and Fisher (2011) -

Syntype workers, queen, U.S.A., California, 1840 [types lost, see Bolton 1977:94].

Myrmica cariniceps Holotype worker, DOMINICAN REPUBLIC, Santo Domingo Island, 19.IV.1850, (A. Salle) (ZSM) [examined].

Myrmica kollari Syntype workers, queen, male, AUSTRIA, Vienna, in hothouses in botanical gardens (NMW) [examined].

Myrmica modesta Syntype workers, INDONESIA, Batjan Island, (A.R. Wallace) (OUMNH) [examined].

Myrmica reticulata Syntype workers, PANAMA, (R.W. Stretch) and UNITED KINGDOM, England, Exeter, botanic gardens, (Parfitt) (BMNH) [examined].

References

References based on Global Ant Biodiversity Informatics

  • Abe T., S. Yamane, and K. Onoyama. Ants collected on the Krakatau Islands 100 years after the great eruptions. Biogeography 14: 65-75.
  • Anh L. N., K. Ogata, and S. Hosoichi. 2010. Ants of agricultural fields in Vietnam (Hymenoptera: Formicidae). Bull. Inst. Trop. Agr., Kyushu Univ. 33: 1-11.
  • Asfiya W., R. Ubaidillah, and Sk. Yamane. 2008. Ants (Hymenoptera: Formicidae) of the Krakataus, and Sebesi and Sebuku islands. Treubia 36: 1-9.
  • Bolton B. 1977. The ant tribe Tetramoriini (Hymenoptera: Formicidae). The genus Tetramorium Mayr in the Oriental and Indo-Australian regions, and in Australia. Bulletin of the British Museum (Natural History). Entomology 36:67-151.
  • Bolton, B. "The ant tribe Tetramoriini (Hymenoptera: Formicinae. The genus Tetramorium Mayr in the Oriental and Indo-Australian regions and in Australia." Bulletin of the British Museum (National History): Entomology series 36, no. 2 (1977): 68-151.
  • Chen P., Y. Su, S. S. Rao, Y. F. Long, and C. H. Du. 2012. Study on the ant diversity in different Illicium verum stands. Journal of West China Forestry Science 41(1): 60-68.
  • Chen Y. Q., Q. Li, Y. L. Chen, Z. X. Lu, X. Y. Zhou. 2011. Ant diversity and bio-indicators in land management of lac insect agroecosystem in Southwestern China. Biodivers. Conserv. 20: 3017-3038.
  • Floren A., W. Wetzel, and M. Staab. 2013. The contribution of canopy species to overall ant diversity (Hymenoptera: Formicidae) in temperate and tropical ecosystems. – Myrmecological News 19: 65-74.
  • Framenau V.W., and M.L. Thomas. 2008. Ants of Christmas Island (Indian Ocean); identification and distribution. Records of the Western Australian Museum 25: 45-85.
  • General D. M., and G. D. Alpert. 2012. A synoptic review of the ant genera (Hymenoptera, Formicidae) of the Philippines. Zookeys 200: 1-111.
  • Gillison A.N. 2000. Above ground biodiversity assesment working group summary report 1996-99: Impact of different land uses on biodiversity and social indicators. ASB Working Group Report, ICRAF, Nairobi, 160pp. http://www.asb.cgiar.org/PDFwebdocs/ASB Biodiversity Report.pdf
  • Guénard B., and R. R. Dunn. 2012. A checklist of the ants of China. Zootaxa 3558: 1-77.
  • Herwina H., N. Nasir, Jumjunidang, and Yaherwandi. 2013. The composition of ant species on banana plants with Banana Bunchy-top Virus (BBTV) symptoms in West Sumatra, Indonesia. Asian Myrmecology 5: 151-161.
  • Hua Li-zhong. 2006. List of Chinese insects Vol. IV. Pages 262-273. Sun Yat-sen university Press, Guangzhou. 539 pages.
  • Huong N. T. T., P. V. Sang, and B. T. Viet. 2015. A preliminary study on diversity of ants (Hymenoptera: Formicidae) at Hon Ba Nature Reserve. Environmental Scientific Conference 7: 614-620.
  • Kadoorie Farm & Botanic Garden, 2002. Report of Rapid Biodiversity Assessments at Qixingkeng Nature Reserve, Southwest Guangdong, 29 April to 1 May and 24 November to 1 December, 1998. South China Biodiversity Survey Report Series (Online Simplified Version): No. 4. KFBG, Hong Kong SAR, ii + 22 pp.
  • Leong C. M., S. F. Shiao, and B. Guenard. 2017. Ants in the city, a preliminary checklist of Formicidae (Hymenoptera) in Macau, one of the most heavily urbanized regions of the world. Asian Myrmecology 9: e009014.
  • Li Q., B. D. Hoffmann, Z. X. Lu, and Y. Q. Chen. 2017. Ants show that the conservation potential of afforestation efforts in Chinese valley-type savanna is dependent upon the afforestation method. Journal of Insect Conservation DOI 10.1007/s10841-017-0005-0
  • Li Q., Z. Lu, Z. Wei, M. Yanyan, and F. Ping. 2015. Communities of ground-dwelling ants in different plantation forest in arid-hot valleys of Jinsha river, Yunnan Province, China. Scientia Silvae Sinicae 51(8): 134-142.
  • Li Qiao, Chen You-qing, Guo Xiao, Duan Yan, Chen Yan-lin, and Xu Zheng-hui. 2007. Diversity of ants in differents habitats in Yuanmou arid-hot valley, Yunnan. Journal of Fujian College of Forestry 27(3): 272-277.
  • Li Z.h. 2006. List of Chinese Insects. Volume 4. Sun Yat-sen University Press
  • Liu X. 2012. Taxonomy, diversity and spatial distribution characters of the ant family Formicidae (Insecta: Hymenoptera) in southeastern Tibet. PhD Thesis 139 pages
  • Liu X., Z. Xu, N. Yu, and C. Zhang. 2016. Distribution patterns of ant species ( Hymenoptera: Formicidae) in Galongla Mountains and Medog Valley of Southeastern Tibet. Scientia Silvae Sinicae 52(11): 88-95.
  • Mathew R. 2003. On Formicidae (Insecta: Hymenoptera) of Nongkhyllem Wild Life Sanctuary, Ri-Bhoi District, Meghalaya. Records of the Zoological Survey of India 101:195-207.
  • Mathew R., and R. N. Tiwari. 2000. Insecta: Hymenoptera: Formicidae. Pp. 251-409 in: Director; Zoological Survey of India (ed.) 2000. Fauna of of Meghalaya. Part 7. [State Fauna Series 4.] Insecta 2000. Calcutta: Zoological Survey of India, 621 pp.
  • Mezger D., and M. Pfeiffer. 2011. Influence of the arrival of Anoplolepis gracilipes (Hymenoptera: Formicidae) on the composition of an ant community in a clearing in Gunung Mulu National Park, Sarawak, Malaysia. Asian Myrmecology 4: 89-98.
  • Mohamed M. 1995. A preliminary list of ants (Hymenoptera: Formicidae) of the Tawau Hills Park, Sabah. In: Ghazally Ismail et al. (eds.), Tawau Hills Park, Sabah, Pelanduk Pub. Pp. 205-213.
  • Ngoc Anh L., K. Ogata, and S. Hosoishi. 2010. Ants of agricultural fields in Vietnam (Hymenoptera: Formicidae). Bull. Inst. Trop. Agr. Kyushu Univ. 33: 1-11.
  • Pfeiffer M., and D. Mezger. 2012. Biodiversity Assessment in Incomplete Inventories: Leaf Litter Ant Communities in Several Types of Bornean Rain Forest. PLoS ONE 7(7): e40729. doi:10.1371/journal.pone.0041000
  • Pfeiffer M.; Mezger, D.; Hosoishi, S.; Bakhtiar, E. Y.; Kohout, R. J. 2011. The Formicidae of Borneo (Insecta: Hymenoptera): a preliminary species list. Asian Myrmecology 4:9-58
  • Rizali A., Clough Y., Buchori D. and Tscharntke T. 2013. Dissimilarity of ant Communities Increases with Precipitation, but not Reduced Land-Use Intensity, in Indonesian Cacao Agroforestry. Diversity. 5: 26-38
  • Rizali A., M. M. Bos, D. Buchori, Sk. Yamane, and C. H. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 15(2): 77-84.
  • Rizali A., M.M. Bos, D. Buchori, Sk. Yamane, C. Hans, and J. Schulze. 2008. Ants in tropical urban habitats: the myrmecofauna in a densely populated area of Bogor, West Java, Indonesia. Hayati Journal of Biosciences 77-84.
  • Rizali A., Y. Clough, D. Buchori, M. L . A. Hosang, M. M. Bos, and T. Tscharntke. 2012. Long-term change of ant community structure in cacao agroforestry landscapes in Indonesia. Insect Conservation and Diversity doi: 10.1111/j.1752-4598.2012.00219.x
  • Song Y., Z. Xu, C. Li, N. Zhang, L. Zhang, H. Jiang, and F. Mo. 2013. An Analysis on the Ant Fauna of the Nangun river Nature Reserve in Yunnan, China. Forest Research 26(6): 773-780.
  • Tang J., Li S., Huang E., Zhang B. and Chen Y. 1995. Hymenoptera: Formicidae (1). Economic Insect Fauna of China 47: 1-133.
  • Terayama M. 2009. A synopsis of the family Formicidae of Taiwan (Insecta: Hymenoptera). Research Bulletin of Kanto Gakuen University. Liberal Arts 17:81-266.
  • Terayama, M. 2009. A synopsis of the family Formicidae of Taiwan (Insecta; Hymenoptera). The Research Bulletin of Kanto Gakuen University 17: 81-266.
  • Torchote P., D. Sitthicharoenchai, and C. Chaisuekul. 2010. Ant species diversity and community composition in three different habitats: mixed deciduous forest, teak plantation and fruit orchard. Tropical Natural History 10(1): 37-51.
  • Trainor C.R. and A.N. Andersen. 2010. The ant fauna of Timor and neighbouring islands: potential bridges between the disjunct faunas of South East Asia and Australia. Australian Journal of Zoology 58: 133-144.
  • Trainor C.R., and A.N. Andersen. 2010. The ant fauna of Timor and neighbouring islands: potential bridges between the disjunct faunas of South East Asia and Australia. Australian Journal of Zoology 58: 133-144.
  • Wang W. R., S. Q. Zhang, and Z. H. Xu. 2012. A faunistic and taxonomic study of ants (Hymenoptera: Formicidae) in Shenzhen Municipality. Journal of Southwest Forestry University 32(1): 64-73.
  • Wang W., S. Zhang, and Z Xu. 2012. Distribution patters of ant species in Shenzhen City. Journal of Southwest Forestry University 32(3): 70-74.
  • Wang W., S. Zhang, and Z. Xu. 2012. Distribution Patterns of Ant Species in Shenzhen City. Journal of Southwest Forestry University 32(3): 69-74.
  • Watanasit S., Noon-anant N., and N. Binnima. 2005. Preliminary survey of ants at a reserve area of Prince of Songkla University, Songkhla Provinces, Southern Thailand. Songklanakarin J. Sci. Technol. 27(1): 39-46.
  • Way M. J., and B. Bolton. 1997. Competition between ants for coconut palm nesting sites. Journal of Natural History 31: 439-455.
  • Wu B., Y. Lu, G. Liang, and L. Zeng. 2010. Influence of the red inported fire ant, Solenopsis invicta Buren (Hymenoptera: Formicidae) on the diversity of ant communities in a newly infested longan orchard and grass areas nearby. Acta Ecologica Sinica 30(8): 2075-2083.
  • Wu B., Y. Lu, L. Zeng, and G. Liang. 2008. Influences of Solenopsis invicta Buren invasion on the native ant communities in different habitats in Guangdong. Chin. J. Appl. Ecol. 19(1): 151-156.
  • Yamane S. 2013. A Review of the ant fauna of the Krakatau Islands, Indonesia. Bull. Kitakyushu Mus. Nat. Hist. Hum. Hist. Ser: A, 11: 1-66
  • Yamane S., T. Itino, and A.R. Nona. 1996. Ground ant fauna in a Bornean dipterocarp forest. Raffles Bulletin of Zoology 44(1): 253-262.
  • Yamane S.; Bui T. V.; Ogata K.; Okido H.; Eguchi K. 2002. Ant fauna of Cuc Phuong National Park, North Vietnam (Hymenoptera: Formicidae). Bulletin of the Institute of Tropical Agriculture Kyushu University 25: 51-62.
  • Yamane S.; Nona, A. R. 1994. Ants from Lambir Hills National Park, Sarawak. Pp. 222-226 in: Inoue, T.; Hamid, A. A. (eds.) 1994. Plant reproductive systems and animal seasonal dynamics. Long-term study of dipterocarp forests in Sarawak. Kyoto: Center for Ecological Research, Kyoto University, vii + 255 pp. 
  • Yamane Sk. 2005. Krakatau in 1982, and the commencement of myrmecological research. The nature and Insects (Konchu to shizen) 40: 27-33.
  • Yamane Sk., T. Itino, and A. Rahman Nona. 1996. Ground ant fauna in a Bornean dipterocarp forest. The Raffles Bulletin of Zoology 44(1): 253-262.
  • Zhang W., G. Liu, P. Zhong, and S. Zhang. 2014. Investigation of Formicidae in Luofushan Mountain. Journal of Huizhou University 34(3): 46-50.
  • Zhou S.-Y. 2001. Ants of Guangxi. Guangxi Normal University Press, Guilin, China, Guilin, China. 255 pp.
  • Zryanin V. A. 2011. An eco-faunistic review of ants (Hymenoptera: Formicidae). In: Structure and functions of soil communities of a monsoon tropical forest (Cat Tien National Park, southern Vietnam) / A.V. Tiunov (Editor). – M.: KMK Scientific Press. 2011. 277 р.101-124.
  • Zryanin V. A., and M. V. Mokrousov. 2015. Contribution to the ant fauna of Lombok Island. Proceedings of the 10th ANeT International Conference, 23-26 October 2015, University of Kelaniya, Sri Lanka. 34