Batrisodes

This is a myrmecophilous genus of Staphylinidae with 89 species. . Adult and immature Batrisodes are generalist predators/ myrmecophiles that inhabit tree holes, forest litter, rotten wood, caves, and are occasionally found in the presence of ants (some species are obligate myrmecophiles). While poorly known at the level of species, Batrisodes is one of the most studied genera within Pselaphinae in terms of biology, largely due to the work of Luigi De Marzo and Orlando Park.

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Batrisodes babaianus Nomura, 2007 g Batrisodes barri Park, 1958 i c g Batrisodes beyeri Schaeffer, 1906 i c g b Batrisodes bistriatus (LeConte, 1849) i c g Batrisodes buqueti (Aubé, 1833) g Batrisodes cartwrighti Sanderson, 1940 i c g Batrisodes cavernosus Park, 1951 i c g Batrisodes cavicornis (Casey, 1897) i c g Batrisodes cicatricosus (Brendel, 1891) i c g Batrisodes clypeonotus (Brendel, 1893) i c g Batrisodes clypeospecus Park, 1960 i c g Batrisodes declivis Casey, 1908 i c g b Batrisodes delaporti (Aubé, 1833) g Batrisodes denticauda (Casey, 1894) i c g Batrisodes denticollis (Casey, 1884) i c g b Batrisodes elysius (Reitter, 1884) g Batrisodes exsculptus (Hampe, 1850) g Batrisodes ferulifer Park, 1960 i c g Batrisodes festinatus Park, 1956 i c g Batrisodes fossicauda (Casey, 1897) i c g Batrisodes foveicornis (Casey, 1887) i c g Batrisodes frontalis (LeConte, 1849) i c g b Batrisodes furcatus (Brendel, 1891) i c g Batrisodes gemmoides Park, 1960 i c g Batrisodes gemmus Park, 1956 i c g\ Batrisodes grubbsi Chandler, 1992 i c g Batrisodes hairstoni Park, 1947 i c g Batrisodes henroti Park, 1956 i c g Batrisodes hubenthali Reitter, 1913 g Batrisodes hubrichti Park, 1958 i c g Batrisodes indistinctus Grigarick and Schuster, 1962 i c g Batrisodes insularis (Baudi di Selve, 1869) g Batrisodes ionae (LeConte, 1849) i c g b Batrisodes jocuvestus Park, 1960 i c g Batrisodes jonesi Park, 1951 i c g Batrisodes juvencus (Brendel, 1865) i c g Batrisodes krekeleri Park, 1960 i c g Batrisodes lineaticollis (Aubé, 1833) i c g b Batrisodes lustrans Casey, 1908 i c g Batrisodes martini Grigarick and Schuster, 1962 i c g Batrisodes masatakai Nomura, 2007 g Batrisodes mendocino (Casey, 1886) i c g b Batrisodes mississippiensis Park, 1956 i c g Batrisodes monticola (Casey, 1886) i c g Batrisodes nebulosus Grigarick and Schuster, 1962 i c g Batrisodes nigricans (LeConte, 1849) i c g Batrisodes obscurus Grigarick and Schuster, 1962 i c g Batrisodes occiduus (Casey, 1886) i c g Batrisodes oculatus (Aubé, 1833) g Batrisodes opacus Grigarick and Schuster, 1962 i c g Batrisodes oro Chandler, 1983 i c g Batrisodes paganettii W.Blattny & C.Blattny, 1916 g Batrisodes pannosus Park, 1960 i c g Batrisodes pogonatus (Saulcy, 1874) g Batrisodes profundus Park, 1956 i c g Batrisodes punctifrons (Casey, 1887) i c g Batrisodes reyesi Chandler, 1992 i c g Batrisodes riparius (Say, 1824) i c g Batrisodes rossi Park, 1947 i c g Batrisodes scabriceps (LeConte, 1849) i c g Batrisodes schaefferi Park, 1947 i c g Batrisodes schaumi (Aubé, 1844) i c g Batrisodes schmitti (Casey, 1897) i c g Batrisodes sinuatifrons (Brendel, 1893) i c g Batrisodes speculum (Casey, 1886) i c g Batrisodes specus Park, 1951 i c g Batrisodes spinifer (Brendel, 1887) i c g Batrisodes spretus (LeConte, 1849) i c g Batrisodes striatus (LeConte, 1849) i c g Batrisodes subterraneus Park, 1951 i c g Batrisodes sulcaticeps Besuchet, 1981 g Batrisodes temporalis (Casey, 1897) i c g Batrisodes texanus Chandler, 1992 i c g b (Coffin Cave mold beetle) Batrisodes tulareanus Casey, 1908 i c g Batrisodes tumoris Park, 1960 i c g Batrisodes uncicornis (Casey, 1897) i c g Batrisodes unisexualis Besuchet, 1988 g Batrisodes valentinei Park, 1951 i c g Batrisodes venustus (Reichenbach, 1816) g Batrisodes venyivi Chandler, 1992 i c g b (helotes mold beetle) Batrisodes virginiae (Casey, 1884) i c Batrisodes wardi Chandler, 2003 i c g Batrisodes yanaorum Chandler, 2003 i c g Batrisodes zephyrinus (Casey, 1886) i c g

List of Batrisodes and their Host Ants

Genus and species	Author and Year	Ant Host	Distribution	Subgenus
Batrisodes abdominalis	Jiang, Ri-Xin & Zi-Wei Yin, 2017	ants	China
Batrisodes acuminatus	(Sharp, 1883)	ants	Japan
Batrisodes acutifrons	Jeannel, 1958	ants	Japan	Excavodes
Batrisodes adnexus	(Hampe, 1863)	ants	EUROPE (AU CR CZ DE FI FR GB GE HU IT PL RO SK SL SV SZ UK YU), RUSSIA
Batrisodes albionicus	(Aubé, 1833)	ants		Empinodes
Batrisodes angustelytratus	Yin, Zi-Wei, Jia-Wei Shen & Li-Zhen Li, 2015	ants	China
Batrisodes angustus	(Sharp, 1874)	ants
Batrisodes antennatus	Schaeffer, 1906	ants	USA (NC,PA,TN,VA)	Babnormodes
Batrisodes aphenogastri	Fall, 1912	Aphaenogaster		Empinodes
Batrisodes asteriscus	(Schaufuss, 1887)	ants
Batrisodes auerbachi	Park, 1956	ants

REFERENCES

• Hampe, C. Ein kleiner Beitrag zur gross-österreichischen Käferfauna. Wiener Entomologische Monatschrift, 7 (8): 285-290. (1863)

• Jeannel, R. Révision des Psélaphides du Japon. Mémoires du Muséum National d'Histoire Naturelle (A: Zoologie), 18 (1): 1-138. (1958).

• Jiang, Ri-Xin & Zi-Wei Yin. Eight new species of Batrisodes Reitter from China (Coleoptera, Staphylinidae, Pselaphinae). ZooKeys, 694: 11–30. (2017).

• Yin, Zi-Wei, Jia-Wei Shen & Li-Zhen Li. New species and new combinations of Asian Batrisodes Reitter (Coleoptera, Staphylinidae, Pselaphinae), and synonymy of Batrisodellus Jeannel with Batrisodes. Deutsche Entomologische Zeitschrift, 62 (1): 45-54. (2015).